Document Detail


Evolution of IESs and scrambling in the actin I gene in hypotrichous ciliates.
MedLine Citation:
PMID:  11742064     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Germ-line (micronuclear) genes in hypotrichous ciliates are interrupted by numerous, short, noncoding, AT-rich segments called internal eliminated segments, or IESs. IESs divide a gene into macronuclear destined segments, or MDSs. IESs are excised from micronuclear genes, and the MDSs are spliced when a micronuclear genome is processed into a macronuclear genome after cell mating. In the micronuclear version of the actin I gene intramolecular recombination between IESs during evolution has put MDSs into a scrambled disorder in some but not all hypotrichs. Studies using rDNA sequences to define phylogenetic relationships among eight hypotrichs suggests that evolution of the micronuclear actin I gene proceeds by successive addition of IESs in earlier diverging species, without MDS scrambling. Continued addition of IESs and recombination among IESs in later diverging species produced actin I genes with scrambled MDSs. Subsequent to MDS scrambling, additional IESs were inserted into the more recently evolved species. Thus, IES insertions and gene scrambling occur in a progressive manner during species evolution to produce micronuclear actin I genes of increasing structural complexity.
Authors:
D J Hogan; E A Hewitt; K E Orr; D M Prescott; K M Müller
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't; Research Support, U.S. Gov't, Non-P.H.S.; Research Support, U.S. Gov't, P.H.S.     Date:  2001-12-11
Journal Detail:
Title:  Proceedings of the National Academy of Sciences of the United States of America     Volume:  98     ISSN:  0027-8424     ISO Abbreviation:  Proc. Natl. Acad. Sci. U.S.A.     Publication Date:  2001 Dec 
Date Detail:
Created Date:  2001-12-25     Completed Date:  2002-01-22     Revised Date:  2009-11-18    
Medline Journal Info:
Nlm Unique ID:  7505876     Medline TA:  Proc Natl Acad Sci U S A     Country:  United States    
Other Details:
Languages:  eng     Pagination:  15101-6     Citation Subset:  IM    
Affiliation:
Department of Molecular, Cellular, and Developmental Biology, University of Colorado, Boulder, CO 80309-034, USA.
Data Bank Information
Bank Name/Acc. No.:
GENBANK/AF188160;  AY044837;  AY044838;  AY044839;  AY046534;  U63566
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MeSH Terms
Descriptor/Qualifier:
Actins / genetics*
Animals
Ciliophora / genetics*
DNA, Ribosomal / genetics
Evolution, Molecular*
Humans
Molecular Sequence Data
Phylogeny
Grant Support
ID/Acronym/Agency:
R01 GM 56161/GM/NIGMS NIH HHS
Chemical
Reg. No./Substance:
0/Actins; 0/DNA, Ribosomal
Comments/Corrections

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