Document Detail


Evidence that Dot-dependent and -independent factors isolate the Legionella pneumophila phagosome from the endocytic network in mouse macrophages.
MedLine Citation:
PMID:  11207624     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Legionella pneumophila survives within macrophages by evading phagosome-lysosome fusion. To determine whether L. pneumophila resides in an intermediate endosomal compartment or is isolated from the endosomal pathway and to investigate what bacterial factors contribute to establishment of its vacuole, we applied a series of fluorescence microscopy assays. The majority of vacuoles, aged 2.5 min to 4 h containing post-exponential phase (PE) L. pneumophila, appeared to be separate from the endosomal pathway, as judged by the absence of transferrin receptor, LAMP-1, cathepsin D and each of four fluorescent probes used to label the endocytic pathway either before or after infection. In contrast, more than 70% of phagosomes that contained Escherichia coli, polystyrene beads, or exponential phase (E) L. pneumophila matured to phagolysosomes, as judged by co-localization with LAMP-1, cathepsin D and fluorescent endosomal probes. Surprisingly, neither bacterial viability nor the putative Dot/Icm transport complex was absolutely required for vacuole isolation; although phagosomes containing either formalin-killed PE wild-type or live PE dotA or dotB mutant L. pneumophila rapidly accumulated LAMP-1, less than 20% acquired lysosomal cathepsin D or fluorescent endosomal probes. Therefore, a Dot-dependent factor(s) isolates the L. pneumophila phagosome from a LAMP-1-containing compartment, and a formalin-resistant Dot-independent activity inhibits vacuolar accumulation of endocytosed material and delivery to the degradative lysosomes.
Authors:
A D Joshi; S Sturgill-Koszycki; M S Swanson
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Publication Detail:
Type:  Journal Article; Research Support, U.S. Gov't, P.H.S.    
Journal Detail:
Title:  Cellular microbiology     Volume:  3     ISSN:  1462-5814     ISO Abbreviation:  Cell. Microbiol.     Publication Date:  2001 Feb 
Date Detail:
Created Date:  2001-02-26     Completed Date:  2001-03-22     Revised Date:  2007-11-14    
Medline Journal Info:
Nlm Unique ID:  100883691     Medline TA:  Cell Microbiol     Country:  England    
Other Details:
Languages:  eng     Pagination:  99-114     Citation Subset:  IM    
Affiliation:
Department of Microbiology and Immunology, The University of Michigan Medical School, Ann Arbor 48109, USA.
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MeSH Terms
Descriptor/Qualifier:
Animals
Antigens, CD / metabolism
Biological Markers
Cathepsin D / metabolism
Endoscopes
Female
Legionella pneumophila / pathogenicity*
Lysosome-Associated Membrane Glycoproteins
Lysosomes
Macrophages / microbiology*
Membrane Glycoproteins / metabolism
Mice
Models, Biological
Molecular Probes
Phagosomes / microbiology*
Receptors, Transferrin / metabolism
Transport Vesicles / microbiology*
Grant Support
ID/Acronym/Agency:
R29AI40694-01BM/AI/NIAID NIH HHS
Chemical
Reg. No./Substance:
0/Antigens, CD; 0/Biological Markers; 0/Lysosome-Associated Membrane Glycoproteins; 0/Membrane Glycoproteins; 0/Molecular Probes; 0/Receptors, Transferrin; EC 3.4.23.5/Cathepsin D

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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