Document Detail


Ethanol-induced methylation of cell cycle genes in neural stem cells.
MedLine Citation:
PMID:  20626555     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Ethanol inhibits the proliferation of neural precursors by altering mitogenic and anti-mitogenic growth factor signaling and can affect global methylation activity in the fetus. We tested the hypothesis that epigenetic modification of specific cell cycle genes underlies the ethanol-induced inhibition of growth factor-regulated cell cycle progression. Monolayer cultures of neural stem cells (NSCs) were treated with fibroblast growth factor 2 or transforming growth factor (TGF) β1 in the absence or presence of ethanol. Ethanol increased the total length of the cell cycle by elongating the amount of time spent in the gap 1 (G1) and synthesis (S) phases of the cell cycle. Ethanol induced the hypermethylation of multiple cell cycle genes associated with the G1/S and gap 2/mitotic phase (G2/M) checkpoints and increased the expression and activity of DNA methyltransferases. These changes were most pronounced in the presence of TGFβ1. Epigenetic alterations paralleled the down-regulation of associated transcripts and other checkpoint-related mRNAs both in vitro (NS-5 cell culture) and in vivo (fetal mouse cortex). Ethanol-induced hypermethylation was accompanied by decreases in the proportion of NSCs expressing associated cell cycle proteins. Thus, ethanol disrupts growth factor-related cell cycle progression by inducing checkpoint restriction at the G1/S transition through a feed-forward system involving the methylation of G2/M regulators.
Authors:
Steven D Hicks; Frank A Middleton; Michael W Miller
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, U.S. Gov't, Non-P.H.S.     Date:  2010-07-27
Journal Detail:
Title:  Journal of neurochemistry     Volume:  114     ISSN:  1471-4159     ISO Abbreviation:  J. Neurochem.     Publication Date:  2010 Sep 
Date Detail:
Created Date:  2010-09-24     Completed Date:  2010-10-19     Revised Date:  2011-03-08    
Medline Journal Info:
Nlm Unique ID:  2985190R     Medline TA:  J Neurochem     Country:  England    
Other Details:
Languages:  eng     Pagination:  1767-80     Citation Subset:  IM    
Copyright Information:
© 2010 The Authors. Journal Compilation © 2010 International Society for Neurochemistry.
Affiliation:
Department of Neuroscience and Physiology, State University of New York - Upstate Medical University, Syracuse, NY 13210, USA.
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MeSH Terms
Descriptor/Qualifier:
Animals
Cell Cycle / drug effects,  genetics*
Cell Cycle Proteins / biosynthesis*,  genetics
Cell Line
Cell Lineage
DNA (Cytosine-5-)-Methyltransferase / biosynthesis
DNA Methylation
Embryonic Stem Cells / drug effects*,  metabolism
Epigenesis, Genetic
Ethanol / pharmacology*
Fibroblast Growth Factor 2 / pharmacology
Gene Expression Regulation
Humans
Kinetics
Mice
Mice, Inbred C57BL
Nerve Tissue / cytology
RNA, Messenger / biosynthesis
Telencephalon / drug effects,  embryology,  metabolism
Transforming Growth Factor beta1 / pharmacology
Grant Support
ID/Acronym/Agency:
AA06916/AA/NIAAA NIH HHS; AA07568/AA/NIAAA NIH HHS; AA16151/AA/NIAAA NIH HHS; AA178231/AA/NIAAA NIH HHS; AA18223/AA/NIAAA NIH HHS; R01 AA006916-26/AA/NIAAA NIH HHS; R37 AA007568-20/AA/NIAAA NIH HHS
Chemical
Reg. No./Substance:
0/Cell Cycle Proteins; 0/RNA, Messenger; 0/Transforming Growth Factor beta1; 103107-01-3/Fibroblast Growth Factor 2; 64-17-5/Ethanol; EC 2.1.1.37/DNA (Cytosine-5-)-Methyltransferase; EC 2.1.1.37/DNA (cytosine-5-)-methyltransferase 1

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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