Document Detail


Endothelial nitric oxide synthase is central to skeletal muscle metabolic regulation and enzymatic signaling during exercise in vivo.
MedLine Citation:
PMID:  20200137     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Endothelial nitric oxide synthase (eNOS) is associated with a number of physiological functions involved in the regulation of metabolism; however, the functional role of eNOS is poorly understood. We tested the hypothesis that eNOS is critical to muscle cell signaling and fuel usage during exercise in vivo, using 16-wk-old catheterized (carotid artery and jugular vein) C57BL/6J mice with wild-type (WT), partial (+/-), or no expression (-/-) of eNOS. Quantitative reductions in eNOS expression ( approximately 40%) elicited many of the phenotypic effects observed in enos(-/-) mice under fasted, sedentary conditions, with expression of oxidative phosphorylation complexes I to V and ATP levels being decreased, and total NOS activity and Ca(2+)/CaM kinase II Thr(286) phosphorylation being increased in skeletal muscle. Despite these alterations, exercise tolerance was markedly impaired in enos(-/-) mice during an acute 30-min bout of exercise. An eNOS-dependent effect was observed with regard to AMP-activated protein kinase signaling and muscle perfusion. Muscle glucose and long-chain fatty acid uptake, and hepatic and skeletal muscle glycogenolysis during the exercise bout was markedly accelerated in enos(-/-) mice compared with enos(+/-) and WT mice. Correspondingly, enos(-/-) mice exhibited hypoglycemia during exercise. Thus, the ablation of eNOS alters a number of physiological processes that result in impaired exercise capacity in vivo. The finding that a partial reduction in eNOS expression is sufficient to induce many of the changes associated with ablation of eNOS has implications for chronic metabolic diseases, such as obesity and insulin resistance, which are associated with reduced eNOS expression.
Authors:
Robert S Lee-Young; Julio E Ayala; Charles F Hunley; Freyja D James; Deanna P Bracy; Li Kang; David H Wasserman
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't     Date:  2010-03-03
Journal Detail:
Title:  American journal of physiology. Regulatory, integrative and comparative physiology     Volume:  298     ISSN:  1522-1490     ISO Abbreviation:  Am. J. Physiol. Regul. Integr. Comp. Physiol.     Publication Date:  2010 May 
Date Detail:
Created Date:  2010-08-17     Completed Date:  2010-09-08     Revised Date:  2011-07-28    
Medline Journal Info:
Nlm Unique ID:  100901230     Medline TA:  Am J Physiol Regul Integr Comp Physiol     Country:  United States    
Other Details:
Languages:  eng     Pagination:  R1399-408     Citation Subset:  IM    
Affiliation:
Department of Molecular Physiology and Biophysics, Vanderbilt University, School of Medicine, 2200 Pierce Ave., Nashville, TN 37232, U.S.A. robert.s.lee-young@vanderbilt.edu
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MeSH Terms
Descriptor/Qualifier:
AMP-Activated Protein Kinases / metabolism
Animals
Body Composition / physiology
Body Weight / physiology
Calorimetry, Indirect
Energy Metabolism / physiology*
Female
Gluconeogenesis / physiology
Glycogen / metabolism
Hypoglycemia / metabolism,  physiopathology
Insulin / blood
Male
Mice
Mice, Inbred C57BL
Mice, Mutant Strains
Mitochondria / physiology
Muscle, Skeletal / blood supply,  enzymology*
Nitric Oxide Synthase Type III / genetics,  metabolism*
Oxidative Phosphorylation
Photoperiod
Physical Conditioning, Animal / physiology
Physical Exertion / physiology*
Pregnancy
Regional Blood Flow / physiology
Signal Transduction / physiology*
Grant Support
ID/Acronym/Agency:
R01 DK-54902/DK/NIDDK NIH HHS; U24 DK-59637/DK/NIDDK NIH HHS
Chemical
Reg. No./Substance:
11061-68-0/Insulin; 9005-79-2/Glycogen; EC 1.14.13.39/Nitric Oxide Synthase Type III; EC 1.14.13.39/Nos3 protein, mouse; EC 2.7.11.1/AMP-Activated Protein Kinases; EC 2.7.11.1/AMPK alpha1 subunit, mouse
Comments/Corrections

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