Document Detail


Effects of Dynein on Microtubule Mechanics and Centrosome Positioning.
MedLine Citation:
PMID:  22013075     Owner:  NLM     Status:  Publisher    
Abstract/OtherAbstract:
In order to determine forces on intracellular microtubules, we measured shape changes of individual microtubules following laser severing. Surprisingly, regions near newly created minus ends increased in curvature following severing, while regions near new microtubule plus ends depolymerized without any observable change in shape. With dynein inhibited, regions near severed minus ends straightened rapidly following severing. These observations suggest that dynein exerts a pulling force on the microtubule which buckles the newly created minus end. Moreover, the lack of any observable straightening suggests that dynein prevents lateral motion of microtubules. To explain these results, we developed a model for intracellular microtubule mechanics which predicts the enhanced buckling at the minus end of a severed microtubule. Our results show that microtubule shapes reflect a dynamic force balance, in which dynein motor and friction forces dominate elastic forces arising from bending moments. A centrosomal array of microtubules subjected to dynein pulling forces and resisted by dynein friction is predicted to center on the experimentally observed timescale, with or without the pushing forces derived from microtubule buckling at the cell periphery.
Authors:
Jun Wu; Gaurav Misra; Robert J Russell; Anthony J C Ladd; Tanmay P Lele; Richard B Dickinson
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Publication Detail:
Type:  JOURNAL ARTICLE     Date:  2011-10-19
Journal Detail:
Title:  Molecular biology of the cell     Volume:  -     ISSN:  1939-4586     ISO Abbreviation:  -     Publication Date:  2011 Oct 
Date Detail:
Created Date:  2011-10-20     Completed Date:  -     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  9201390     Medline TA:  Mol Biol Cell     Country:  -    
Other Details:
Languages:  ENG     Pagination:  -     Citation Subset:  -    
Affiliation:
*Department of Chemical Engineering, University of Florida, Gainesville FL 32611.
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