Document Detail


Drosophila CAF-1 regulates HP1-mediated epigenetic silencing and pericentric heterochromatin stability.
MedLine Citation:
PMID:  20663913     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Chromatin assembly factor 1 (CAF-1) was initially characterized as a histone deliver in the process of DNA-replication-coupled chromatin assembly in eukaryotic cells. Here, we report that CAF-1 p180, the largest subunit of Drosophila CAF-1, participates in the process of heterochromatin formation and functions to maintain pericentric heterochromatin stability. We provide evidence that Drosophila CAF-1 p180 plays a role in both classes of position effect variegation (PEV) and in the expression of heterochromatic genes. A decrease in the expression of Drosophila CAF-1 p180 leads to a decrease in both H3K9 methylation at pericentric heterochromatin regions and the recruitment of heterochromatin protein 1 (HP1) to the chromocenter of the polytene chromosomes. The artificial targeting of HP1 to a euchromatin location leads to the enrichment of Drosophila CAF-1 p180 at this ectopic heterochromatin, suggesting the mutual recruitment of HP1 and CAF-1 p180. We also show that the spreading of heterochromatin is compromised in flies that have reduced CAF-1 p180. Furthermore, reduced CAF-1 p180 causes a defect in the dynamics of heterochromatic markers in early Drosophila embryos. Together, these findings suggest that Drosophila CAF-1 p180 is an essential factor in the epigenetic control of heterochromatin formation and/or maintenance.
Authors:
Hai Huang; Zhongsheng Yu; Shuaiqi Zhang; Xuehong Liang; Jianming Chen; Changqing Li; Jun Ma; Renjie Jiao
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2010-07-27
Journal Detail:
Title:  Journal of cell science     Volume:  123     ISSN:  1477-9137     ISO Abbreviation:  J. Cell. Sci.     Publication Date:  2010 Aug 
Date Detail:
Created Date:  2010-08-05     Completed Date:  2010-12-02     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  0052457     Medline TA:  J Cell Sci     Country:  England    
Other Details:
Languages:  eng     Pagination:  2853-61     Citation Subset:  IM    
Affiliation:
State Key Laboratory of Brain and Cognitive Science, Institute of Biophysics, Chinese Academy of Sciences, Beijing 100101, China.
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MeSH Terms
Descriptor/Qualifier:
Animals
Drosophila / genetics*
Drosophila Proteins / genetics*,  metabolism
Epigenesis, Genetic
Gene Expression Regulation
Gene Silencing
Heterochromatin / genetics*,  metabolism
Repressor Proteins / genetics,  metabolism
Retinoblastoma-Binding Protein 4 / genetics*,  metabolism
Chemical
Reg. No./Substance:
0/CAF1 protein, Drosophila; 0/Drosophila Proteins; 0/Heterochromatin; 0/Repressor Proteins; 0/Retinoblastoma-Binding Protein 4

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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