Document Detail


A DNA segment conferring stable maintenance on R6K gamma-origin core replicons.
MedLine Citation:
PMID:  7592407     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
The plasmid R6K gamma origin consists of two adjacent modules, the enhancer and the core, and requires R6K initiator protein pi for replication. While the core alone can replicate at a low level of wild-type pi protein, we show here that host cells do not stably maintain core plasmids. The presence of the enhancer segment confers stable inheritance on core plasmids without a significant change in average plasmid copy number. Deletions and site-directed mutagenesis indicated that the stability of core plasmids is not mediated by binding sites or consensus sequences in the enhancer for DnaA, pi protein, gyrase, Fis, or Dcm methylase. Proper segregation of core plasmids requires only the R6K stb or stability-related region, which includes the 20-bp segment of the 100-bp enhancer adjacent to the core. The use of the pi 116 mutant protein, which increases plasmid copy number fourfold, does not stabilize core plasmids lacking the enhancer. We also show that at an elevated level of wild-type pi, the gamma-origin plasmid is unstable, even in the presence of the enhancer. We discuss the differences and similarities between the R6K stability system and those found in other plasmids.
Authors:
F Wu; I Levchenko; M Filutowicz
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Publication Detail:
Type:  Journal Article; Research Support, U.S. Gov't, P.H.S.    
Journal Detail:
Title:  Journal of bacteriology     Volume:  177     ISSN:  0021-9193     ISO Abbreviation:  J. Bacteriol.     Publication Date:  1995 Nov 
Date Detail:
Created Date:  1995-12-19     Completed Date:  1995-12-19     Revised Date:  2009-11-18    
Medline Journal Info:
Nlm Unique ID:  2985120R     Medline TA:  J Bacteriol     Country:  UNITED STATES    
Other Details:
Languages:  eng     Pagination:  6338-45     Citation Subset:  IM    
Affiliation:
Department of Bacteriology, University of Wisconsin, Madison 53706, USA.
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MeSH Terms
Descriptor/Qualifier:
Bacterial Proteins / metabolism
Base Sequence
DNA Helicases*
DNA, Bacterial / genetics,  metabolism
DNA-Binding Proteins / metabolism
Enhancer Elements, Genetic / genetics*
Escherichia coli / genetics
Gene Dosage
Molecular Sequence Data
Mutagenesis
Peptide Initiation Factors / metabolism
R Factors / genetics,  metabolism*
Replication Origin / genetics*
Trans-Activators / metabolism
Grant Support
ID/Acronym/Agency:
GM40314/GM/NIGMS NIH HHS
Chemical
Reg. No./Substance:
0/Bacterial Proteins; 0/DNA, Bacterial; 0/DNA-Binding Proteins; 0/Peptide Initiation Factors; 0/Trans-Activators; 0/replication initiator protein; EC 3.6.1.-/DNA Helicases
Comments/Corrections

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