Document Detail


Coupling prokaryotic cell fate and division control with a bifunctional and oscillating oxidoreductase homolog.
MedLine Citation:
PMID:  20152180     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
NAD(H)-binding proteins play important roles in cell-cycle and developmental signaling in eukaryotes. We identified a bifunctional NAD(H)-binding regulator (KidO) that integrates cell-fate signaling with cytokinesis in the bacterium Caulobacter crescentus. KidO stimulates the DivJ kinase and directly acts on the cytokinetic tubulin, FtsZ, to tune cytokinesis with the cell cycle. At the G1-->S transition, DivJ concomitantly signals the ClpXP-dependent degradation of KidO and CtrA, a cell-cycle transcriptional regulator/DNA replication inhibitor. This proteolytic event directs KidO and CtrA into oscillatory cell-cycle abundance patterns that coordinately license replication and cytokinesis. KidO resembles NAD(P)H-dependent oxidoreductases, and conserved residues in the KidO NAD(H)-binding pocket are critical for regulation of FtsZ, but not for DivJ. Since NADPH-dependent regulation by a KidO-like oxidoreductase also occurs in humans, organisms from two domains of life exploit the enzymatic fold of an ancestral oxidoreductase potentially to coordinate cellular or developmental activities with the availability of the metabolic currency, NAD(P)H.
Authors:
Sunish Kumar Radhakrishnan; Sean Pritchard; Patrick H Viollier
Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't; Research Support, U.S. Gov't, Non-P.H.S.    
Journal Detail:
Title:  Developmental cell     Volume:  18     ISSN:  1878-1551     ISO Abbreviation:  Dev. Cell     Publication Date:  2010 Jan 
Date Detail:
Created Date:  2010-02-15     Completed Date:  2010-03-19     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  101120028     Medline TA:  Dev Cell     Country:  United States    
Other Details:
Languages:  eng     Pagination:  90-101     Citation Subset:  IM    
Copyright Information:
(c) 2010 Elsevier Inc. All rights reserved.
Affiliation:
Department of Molecular Biology and Microbiology, School of Medicine, Case Western Reserve University, Cleveland, OH 44106, USA.
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MeSH Terms
Descriptor/Qualifier:
Bacterial Proteins / metabolism
Biological Clocks / physiology*
Carrier Proteins / metabolism
Catalytic Domain / physiology
Caulobacter crescentus / enzymology*,  genetics
Cell Cycle / physiology
Cell Cycle Proteins / genetics,  metabolism
Cell Division / physiology*
Cell Lineage / physiology*
Cytoskeletal Proteins / metabolism
NAD / metabolism
Oxidation-Reduction
Oxidoreductases / genetics,  metabolism*
Phosphotransferases / metabolism
Prokaryotic Cells / enzymology
Tubulin / metabolism
Grant Support
ID/Acronym/Agency:
SIGRR016789-01A1//PHS HHS
Chemical
Reg. No./Substance:
0/Bacterial Proteins; 0/Carrier Proteins; 0/Cell Cycle Proteins; 0/Cytoskeletal Proteins; 0/FtsZ protein, Bacteria; 0/Tubulin; 53-84-9/NAD; EC 1.-/Oxidoreductases; EC 2.7.-/Phosphotransferases

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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