Document Detail


Control of cell proliferation, organ growth, and DNA damage response operate independently of dephosphorylation of the Arabidopsis Cdk1 homolog CDKA;1.
MedLine Citation:
PMID:  19948791     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Entry into mitosis is universally controlled by cyclin-dependent kinases (CDKs). A key regulatory event in metazoans and fission yeast is CDK activation by the removal of inhibitory phosphate groups in the ATP binding pocket catalyzed by Cdc25 phosphatases. In contrast with other multicellular organisms, we show here that in the flowering plant Arabidopsis thaliana, cell cycle control does not depend on sudden changes in the phosphorylation pattern of the PSTAIRE-containing Cdk1 homolog CDKA;1. Consistently, we found that neither mutants in a previously identified CDC25 candidate gene nor plants in which it is overexpressed display cell cycle defects. Inhibitory phosphorylation of CDKs is also the key event in metazoans to arrest cell cycle progression upon DNA damage. However, we show here that the DNA damage checkpoint in Arabidopsis can also operate independently of the phosphorylation of CDKA;1. These observations reveal a surprising degree of divergence in the circuitry of highly conserved core cell cycle regulators in multicellular organisms. Based on biomathematical simulations, we propose a plant-specific model of how progression through the cell cycle could be wired in Arabidopsis.
Authors:
Nico Dissmeyer; Annika K Weimer; Stefan Pusch; Kristof De Schutter; Claire Lessa Alvim Kamei; Moritz K Nowack; Bela Novak; Gui-Lan Duan; Yong-Guan Zhu; Lieven De Veylder; Arp Schnittger
Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2009-11-30
Journal Detail:
Title:  The Plant cell     Volume:  21     ISSN:  1532-298X     ISO Abbreviation:  Plant Cell     Publication Date:  2009 Nov 
Date Detail:
Created Date:  2010-01-01     Completed Date:  2010-03-22     Revised Date:  2011-09-08    
Medline Journal Info:
Nlm Unique ID:  9208688     Medline TA:  Plant Cell     Country:  United States    
Other Details:
Languages:  eng     Pagination:  3641-54     Citation Subset:  IM    
Affiliation:
Unigruppe am Max-Planck-Institut für Züchtungsforschung, Max-Delbrück-Laboratorium, Lehrstuhl für Botanik III, Universität zu Köln, Köln, Germany. diss@mpiz-koeln.mpg.de
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MeSH Terms
Descriptor/Qualifier:
Arabidopsis / genetics*,  growth & development,  metabolism
Arabidopsis Proteins / genetics*,  metabolism
Binding Sites / genetics
CDC2 Protein Kinase / genetics,  metabolism
Cell Cycle / genetics
Cell Cycle Proteins / genetics*,  metabolism
Cell Proliferation*
Cyclin-Dependent Kinases / genetics*,  metabolism
DNA Damage / genetics*
DNA Repair / genetics*
Evolution, Molecular
Genes, cdc / physiology
Mathematics
Mitosis / genetics
Phosphorylation
Plant Structures / genetics,  growth & development,  metabolism
cdc25 Phosphatases / genetics,  metabolism
Chemical
Reg. No./Substance:
0/Arabidopsis Proteins; 0/CDKA1 protein, Arabidopsis; 0/Cell Cycle Proteins; EC 2.7.11.22/CDC2 Protein Kinase; EC 2.7.11.22/Cyclin-Dependent Kinases; EC 3.1.3.48/cdc25 Phosphatases

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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