Document Detail


Cognate interactions between helper T cells and B cells. III. Contact-dependent, lymphokine-independent induction of B cell cycle entry by activated helper T cells.
MedLine Citation:
PMID:  2476483     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
An Ag-specific, IL-2-dependent Th clone induced the growth of B cells in a class II-restricted, Ag-specific, IL-2-dependent manner. The formation of stable Th-3.1-B cell conjugates was restricted by Ag and class II MHC. After activation of Th-3.1 by insolubilized anti-T3 (Th-3.1T3), Th-3.1T3 induced the growth of B cells in a class II unrestricted, Ag nonspecific manner. The formation of stable conjugates between Th-3.1T3 and B cells was also class II unrestricted and Ag nonspecific. Although the interaction of Th-3.1T3 and B cells was class II unrestricted, the interaction was inhibited by the combination of anti-IA and anti-IE mAb. This suggested that monomorphic domains of class II MHC molecules were involved in Th-3.1T3-B cell interaction. Fixed Th-3.1T3 but not fixed resting Th-3.1 induced B cell cycle entry, as measured by an increase in B cell RNA synthesis. Trypsin-treatment of Th-3.1T3 before fixation reduced their ability to activate B cells, indicating that cell surface proteins on Th-3.1T3 were required for enhanced B cell RNA synthesis. Anti-IL-4, anti-IL-2R, or anti-IFN-gamma did not affect the ability of Th-3.1T3 to induce heightened B cell RNA synthesis. Progression into S phase by B cells activated with fixed Th-3.1T3 was supported by the addition of soluble factors. When stimulated with fixed Th-3.1T3, EL4 supernatant (SN) enhanced B cell DNA synthesis. Depletion of IL-4, but not IL-2, from EL4 SN ablated its supportive capabilities. IL-4 alone was completely ineffective in supporting entry into S phase. Therefore, IL-4 and another activity(ies) in EL4 SN were necessary for B cell cycle progression into S phase. Taken together, these data suggest that after Th activation, Th cell surface proteins are expressed that mediate the binding of Th to B cells via recognition of nonpolymorphic domains of class II MHC molecules. Contact of Th-3.1T3 with B cells, not lymphokines, results in the entry of B cells into the cell cycle and heightened B cell lymphokine responsiveness. The addition of exogenous lymphokines supports the progression of Th-3.1T3-activated B cells into S phase.
Authors:
R J Noelle; J McCann; L Marshall; W C Bartlett
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Publication Detail:
Type:  Journal Article; Research Support, U.S. Gov't, P.H.S.    
Journal Detail:
Title:  Journal of immunology (Baltimore, Md. : 1950)     Volume:  143     ISSN:  0022-1767     ISO Abbreviation:  J. Immunol.     Publication Date:  1989 Sep 
Date Detail:
Created Date:  1989-10-18     Completed Date:  1989-10-18     Revised Date:  2008-11-21    
Medline Journal Info:
Nlm Unique ID:  2985117R     Medline TA:  J Immunol     Country:  UNITED STATES    
Other Details:
Languages:  eng     Pagination:  1807-14     Citation Subset:  AIM; IM    
Affiliation:
Department of Microbiology, Dartmouth Medical School, Hanover, NH 03755.
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MeSH Terms
Descriptor/Qualifier:
Animals
Antibodies, Monoclonal / physiology
Antigens, CD3
Antigens, Differentiation, T-Lymphocyte / immunology
B-Lymphocytes / immunology*,  physiology
Cell Communication*
Cell Cycle*
DNA / biosynthesis
Epitopes
Female
Fixatives
Histocompatibility Antigens Class II
Interferon-gamma / physiology
Interleukin-4
Interleukins / physiology
Interphase
Lymphocyte Activation*
Lymphokines / physiology*
Mice
Mice, Inbred C57BL
Mice, Inbred DBA
Receptors, Antigen, T-Cell / immunology
Receptors, Interleukin-2 / physiology
T-Lymphocytes, Helper-Inducer / immunology*,  physiology
Grant Support
ID/Acronym/Agency:
GM-36814/GM/NIGMS NIH HHS; GM37767/GM/NIGMS NIH HHS
Chemical
Reg. No./Substance:
0/Antibodies, Monoclonal; 0/Antigens, CD3; 0/Antigens, Differentiation, T-Lymphocyte; 0/Epitopes; 0/Fixatives; 0/Histocompatibility Antigens Class II; 0/Interleukins; 0/Lymphokines; 0/Receptors, Antigen, T-Cell; 0/Receptors, Interleukin-2; 207137-56-2/Interleukin-4; 82115-62-6/Interferon-gamma; 9007-49-2/DNA

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