Document Detail


Cleavage of the adaptor protein TRIF by enterovirus 71 3C inhibits antiviral responses mediated by Toll-like receptor 3.
MedLine Citation:
PMID:  21697485     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Enterovirus 71 (EV71) causes hand-foot-and-mouth disease and neurological complications in young children. Although the underlying mechanisms remain obscure, impaired or aberrant immunity is thought to play a role. In infected cells, EV71 suppresses type I interferon responses mediated by retinoid acid-inducible gene I (RIG-I). This involves the EV71 3C protein, which disrupts the formation of a functional RIG-I complex. In the present study, we report that EV71 inhibits the induction of innate immunity by Toll-like receptor 3 (TLR3) via a distinct mechanism. In HeLa cells stimulated with poly(I · C), EV71 inactivates interferon regulatory factor 3 and drastically suppresses interferon-stimulated gene expression. Notably, EV71 specifically downregulates a TRIF, TIR domain-containing adaptor inducing beta interferon (IFN-β). When expressed alone in mammalian cells, EV71 3C is capable of exhibiting these activities. EV71 3C associates with and induces TRIF cleavage in the presence of Z-VAD-FMK, a caspase inhibitor. TRIF cleavage depends on its amino acid pair Q312-S313, which resembles a proteolytic site of picornavirus 3C proteases. Further, site-specific 3C mutants with a defective protease activity bind TRIF but fail to mediate TRIF cleavage. Consequently, these 3C mutants are unable to inhibit NF-κB and IFN-β promoter activation. TRIF cleavage mediated by EV71 may be a mechanism to impair type I IFN production in response to Toll-like receptor 3 (TLR3) activation.
Authors:
Xiaobo Lei; Zhenmin Sun; Xinlei Liu; Qi Jin; Bin He; Jianwei Wang
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't     Date:  2011-06-22
Journal Detail:
Title:  Journal of virology     Volume:  85     ISSN:  1098-5514     ISO Abbreviation:  J. Virol.     Publication Date:  2011 Sep 
Date Detail:
Created Date:  2011-08-09     Completed Date:  2011-10-04     Revised Date:  2012-03-01    
Medline Journal Info:
Nlm Unique ID:  0113724     Medline TA:  J Virol     Country:  United States    
Other Details:
Languages:  eng     Pagination:  8811-8     Citation Subset:  IM    
Affiliation:
State Key Laboratory of Molecular Virology and Genetic Engineering, Institute of Pathogen Biology, Chinese Academy of Medical Sciences and Peking Union Medical College, Beijing 100730, People's Republic of China.
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MeSH Terms
Descriptor/Qualifier:
Adaptor Proteins, Vesicular Transport / metabolism*
Cysteine Endopeptidases / genetics,  metabolism*
Enterovirus A, Human / immunology*,  pathogenicity*
HeLa Cells
Host-Pathogen Interactions*
Humans
Mutagenesis, Site-Directed
Mutant Proteins / genetics,  metabolism
Protein Binding
Protein Interaction Mapping
Toll-Like Receptor 3 / immunology*
Viral Proteins / genetics,  metabolism*
Grant Support
ID/Acronym/Agency:
AI092230/AI/NIAID NIH HHS
Chemical
Reg. No./Substance:
0/Adaptor Proteins, Vesicular Transport; 0/Mutant Proteins; 0/TICAM1 protein, human; 0/TLR3 protein, human; 0/Toll-Like Receptor 3; 0/Viral Proteins; EC 3.4.22.-/Cysteine Endopeptidases; EC 3.4.22.28/3C proteases

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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