Document Detail


Chemokine-induced Zap70 kinase-mediated dissociation of the Vav1-talin complex activates alpha4beta1 integrin for T cell adhesion.
MedLine Citation:
PMID:  20005136     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Lymphocyte integrins mediate cell arrest on endothelium during immune surveillance after activation by chemokine-stimulated inside-out signals. Here we show that a Vav1-talin complex in T cells is a key target for chemokine-triggered inside-out signaling leading to integrin alpha4beta1 activation. Thus, Vav1 dissociation from talin was required to generate high-affinity alpha4beta1 conformations. Assembly of the Vav1-talin complex required PtdIns(4,5)P(2), which was provided by talin-bound phosphatidylinositol phosphate kinase Igamma. Chemokine-promoted Vav1 dissociation from talin followed an initial increase in talin binding to alpha4beta1. This process was dependent on ZAP-70, which binds to and phosphorylates Vav1 in the complex, leading to further alpha4beta1 activation and cell adhesion strengthening. Moreover, Vav1-talin dissociation was needed for Rac1 activation, thus indicating that alpha4beta1 and Rac1 activation can be coupled by chemokine-stimulated ZAP-70 function. Our data suggest that Vav1 might function as a repressive adaptor of talin that must dissociate from alpha4beta1-talin complexes for efficient integrin activation.
Authors:
David Garc?a-Bernal; Marisa Parmo-Caba?as; Ana Dios-Esponera; Rafael Samaniego; Dolores Hern?n-P de la Ossa; Joaquin Teixid?
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2009-12-10
Journal Detail:
Title:  Immunity     Volume:  31     ISSN:  1097-4180     ISO Abbreviation:  Immunity     Publication Date:  2009 Dec 
Date Detail:
Created Date:  2010-01-12     Completed Date:  2010-02-09     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  9432918     Medline TA:  Immunity     Country:  United States    
Other Details:
Languages:  eng     Pagination:  953-64     Citation Subset:  IM    
Copyright Information:
Copyright 2009 Elsevier Inc. All rights reserved.
Affiliation:
Cellular and Molecular Medicine Program, Centro de Investigaciones Biol?gicas (CSIC), 28040 Madrid, Spain.
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MeSH Terms
Descriptor/Qualifier:
Cell Adhesion
Cell Line, Tumor
Cells, Cultured
Chemokine CXCL12 / pharmacology
Humans
Integrin alpha4beta1 / metabolism*
Phosphotransferases / metabolism
Proto-Oncogene Proteins c-vav / drug effects,  metabolism*
RNA Interference
Signal Transduction
T-Lymphocytes / drug effects,  immunology*
Talin / drug effects,  metabolism*
Transfection
Vascular Cell Adhesion Molecule-1 / metabolism
ZAP-70 Protein-Tyrosine Kinase / metabolism*
rac1 GTP-Binding Protein / metabolism
Chemical
Reg. No./Substance:
0/Chemokine CXCL12; 0/Integrin alpha4beta1; 0/Proto-Oncogene Proteins c-vav; 0/Talin; 0/VAV1 protein, human; 0/Vascular Cell Adhesion Molecule-1; EC 2.7.-/Phosphotransferases; EC 2.7.1.112/ZAP70 protein, human; EC 2.7.10.2/ZAP-70 Protein-Tyrosine Kinase; EC 3.6.5.2/rac1 GTP-Binding Protein

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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