Document Detail


Changes in plant mitochondrial electron transport alter cellular levels of reactive oxygen species and susceptibility to cell death signaling molecules.
MedLine Citation:
PMID:  17012741     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Transgenic tobacco (Nicotiana tabacum) lacking mitochondrial alternative oxidase (AOX) have been compared with wild-type (Wt) tobacco using two different systems, either suspension cell cultures or leaves. In both systems, a lack of AOX was accompanied by an increase in some anti-oxidant defenses, consistent with the hypothesis that a lack of AOX increases the mitochondrial generation of reactive oxygen species (ROS). In most cases, this increase in anti-oxidant defenses could more than offset the presumed increased rate of ROS generation, resulting paradoxically in a lower steady-state level of ROS than was found in Wt leaves or suspension cells. We also found that the amount of cell death induced by salicylic acid or nitric oxide correlated strongly with the level of ROS (irrespective of the level of AOX), while death induced by azide was dependent upon the presence or absence of AOX. These results suggest that susceptibility to cell death by signaling molecules (salicylic acid and nitric oxide) is dependent upon the steady-state cellular level of ROS and that AOX levels clearly contribute to this steady state, perhaps by influencing the rate of mitochondrial-generated ROS and hence the cellular level of anti-oxidant defenses.
Authors:
Sasan Amirsadeghi; Christine A Robson; Allison E McDonald; Greg C Vanlerberghe
Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2006-09-30
Journal Detail:
Title:  Plant & cell physiology     Volume:  47     ISSN:  0032-0781     ISO Abbreviation:  Plant Cell Physiol.     Publication Date:  2006 Nov 
Date Detail:
Created Date:  2006-11-23     Completed Date:  2007-02-13     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  9430925     Medline TA:  Plant Cell Physiol     Country:  Japan    
Other Details:
Languages:  eng     Pagination:  1509-19     Citation Subset:  IM    
Affiliation:
Department of Life Sciences, University of Toronto Scarborough, 1265 Military Trail, Toronto, ON, Canada M1C1A4.
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MeSH Terms
Descriptor/Qualifier:
Antioxidants / metabolism
Cell Death / drug effects
Electron Transport / drug effects*
Electron Transport Complex IV / metabolism
Gene Expression Regulation, Plant / drug effects
Genes, Plant
Mitochondria / drug effects,  metabolism*
Nitric Oxide / pharmacology*
Oxidoreductases / deficiency
Plant Leaves / drug effects
Plant Proteins / metabolism
Plants, Genetically Modified
RNA, Messenger / genetics,  metabolism
Reactive Oxygen Species / metabolism*
Salicylic Acid / pharmacology*
Tobacco / cytology,  drug effects,  metabolism*
Up-Regulation / drug effects
Chemical
Reg. No./Substance:
0/Antioxidants; 0/Plant Proteins; 0/RNA, Messenger; 0/Reactive Oxygen Species; 10102-43-9/Nitric Oxide; 69-72-7/Salicylic Acid; EC 1.-/Oxidoreductases; EC 1.-/alternative oxidase; EC 1.9.3.1/Electron Transport Complex IV

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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