Document Detail


Changes in cyclin dependent kinase expression and activity accompanying lens fiber cell differentiation.
MedLine Citation:
PMID:  10620399     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Previous studies from this laboratory have shown that differentiating lens fiber cells contain two active cyclin dependent kinases (Cdks), Cdk1 and Cdk5. The present study was undertaken to explore the expression and regulation of six additional members of the Cdk family (Cdk2, Cdk3, Cdk4, Cdk6, Cdk7 and Cdk8) during lens differentiation. Differentiating lens fiber cells were separated from lens epithelial cells by microdissection of developing rat lenses [embryonic day 16 (E16) to postnatal day 8 (P8)] and Cdk expression was assessed by RT-PCR and immunoblotting. Two Cdks (Cdk3 and Cdk6) were not expressed in lens fiber cells or epithelial cells during this developmental period. In the lens epithelium, we detected proteins and mRNAs corresponding to all other Cdks examined (Cdk2, Cdk4, Cdk7, Cdk8) throughout this developmental period. Epithelial cells showed significant Cdk2 activity, which decreased with developmental age, but no significant activity was detected for Cdk4, Cdk7, or Cdk8. Fiber cells contained all four Cdk proteins and the corresponding Cdk mRNAs except for Cdk2 mRNA. None of the Cdks examined showed significant kinase activity in fiber cells. Immunoprecipitates of Cdk2 and Cdk4 from fiber cells contained p57(kip2), supporting the view that this Cdk inhibitor blocks the activity of these Cdks in lens fibers. In contrast, p57(kip2)did not co-immunoprecipitate with Cdk5 from lens fibers. These findings suggest that the differential affinity of p57(kip2)for members of the Cdk family may provide a mechanism for specific regulation of individual Cdks during fiber cell differentiation.
Authors:
C Y Gao; A M Rampalli; H C Cai; H Y He; P S Zelenka
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Publication Detail:
Type:  Journal Article    
Journal Detail:
Title:  Experimental eye research     Volume:  69     ISSN:  0014-4835     ISO Abbreviation:  Exp. Eye Res.     Publication Date:  1999 Dec 
Date Detail:
Created Date:  2000-02-24     Completed Date:  2000-02-24     Revised Date:  2012-06-25    
Medline Journal Info:
Nlm Unique ID:  0370707     Medline TA:  Exp Eye Res     Country:  ENGLAND    
Other Details:
Languages:  eng     Pagination:  695-703     Citation Subset:  IM    
Copyright Information:
Copyright 1999 Academic Press.
Affiliation:
Laboratory of Molecular and Developmental Biology, NEI/NIH, Bethesda, MD, 20892-2730, USA.
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MeSH Terms
Descriptor/Qualifier:
Animals
CDC2-CDC28 Kinases*
Cyclin-Dependent Kinase 2
Cyclin-Dependent Kinase 4
Cyclin-Dependent Kinases / genetics*,  metabolism
DNA Primers
Electrophoresis, Polyacrylamide Gel
Embryonic Induction
Epithelial Cells / metabolism
Gene Expression
Immunoblotting
Lens, Crystalline / cytology,  embryology*,  metabolism
Precipitin Tests
Protein-Serine-Threonine Kinases / genetics*,  metabolism
Proto-Oncogene Proteins*
Rats
Rats, Wistar
Reverse Transcriptase Polymerase Chain Reaction
Chemical
Reg. No./Substance:
0/DNA Primers; 0/Proto-Oncogene Proteins; EC 2.7.11.1/Protein-Serine-Threonine Kinases; EC 2.7.11.22/CDC2-CDC28 Kinases; EC 2.7.11.22/Cdk2 protein, rat; EC 2.7.11.22/Cdk4 protein, rat; EC 2.7.11.22/Cyclin-Dependent Kinase 2; EC 2.7.11.22/Cyclin-Dependent Kinase 4; EC 2.7.11.22/Cyclin-Dependent Kinases; EC 2.7.11.22/cyclin-dependent kinase-activating kinase

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