Document Detail


Cell ingression and apical shape oscillations during dorsal closure in Drosophila.
MedLine Citation:
PMID:  22404919     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Programmed patterns of gene expression, cell-cell signaling, and cellular forces cause morphogenic movements during dorsal closure. We investigated the apical cell-shape changes that characterize amnioserosa cells during dorsal closure in Drosophila embryos with in vivo imaging of green-fluorescent-protein-labeled DE-cadherin. Time-lapsed, confocal images were assessed with a novel segmentation algorithm, Fourier analysis, and kinematic and dynamical modeling. We found two generic processes, reversible oscillations in apical cross-sectional area and cell ingression characterized by persistent loss of apical area. We quantified a time-dependent, spatially-averaged sum of intracellular and intercellular forces acting on each cell's apical belt of DE-cadherin. We observed that a substantial fraction of amnioserosa cells ingress near the leading edges of lateral epidermis, consistent with the view that ingression can be regulated by leading-edge cells. This is in addition to previously observed ingression processes associated with zipping and apoptosis. Although there is cell-to-cell variability in the maximum rate for decreasing apical area (0.3-9.5 μm(2)/min), the rate for completing ingression is remarkably constant (0.83 cells/min, r(2) > 0.99). We propose that this constant ingression rate contributes to the spatiotemporal regularity of mechanical stress exerted by the amnioserosa on each leading edge during closure.
Authors:
Adam Sokolow; Yusuke Toyama; Daniel P Kiehart; Glenn S Edwards
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't     Date:  2012-03-06
Journal Detail:
Title:  Biophysical journal     Volume:  102     ISSN:  1542-0086     ISO Abbreviation:  Biophys. J.     Publication Date:  2012 Mar 
Date Detail:
Created Date:  2012-03-12     Completed Date:  2012-07-05     Revised Date:  2013-04-16    
Medline Journal Info:
Nlm Unique ID:  0370626     Medline TA:  Biophys J     Country:  United States    
Other Details:
Languages:  eng     Pagination:  969-79     Citation Subset:  IM    
Copyright Information:
Copyright © 2012 Biophysical Society. Published by Elsevier Inc. All rights reserved.
Affiliation:
Physics Department, Duke University, Durham, North Carolina, USA.
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MeSH Terms
Descriptor/Qualifier:
Animals
Cadherins / metabolism
Cell Polarity
Cell Shape*
Drosophila melanogaster / cytology,  embryology*
Embryo, Nonmammalian / cytology*,  metabolism
Embryonic Development*
Models, Biological*
Time Factors
Grant Support
ID/Acronym/Agency:
GM 33830/GM/NIGMS NIH HHS
Chemical
Reg. No./Substance:
0/Cadherins
Comments/Corrections

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