Document Detail


The CcrM DNA methyltransferase of Agrobacterium tumefaciens is essential, and its activity is cell cycle regulated.
MedLine Citation:
PMID:  11325934     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
DNA methylation is now recognized as a regulator of multiple bacterial cellular processes. CcrM is a DNA adenine methyltransferase found in the alpha subdivision of the proteobacteria. Like the Dam enzyme, which is found primarily in Escherichia coli and other gamma proteobacteria, it does not appear to be part of a DNA restriction-modification system. The CcrM homolog of Agrobacterium tumefaciens was found to be essential for viability. Overexpression of CcrM is associated with significant abnormalities of cell morphology and DNA ploidy. Mapping of the transcriptional start site revealed a conserved binding motif for the global response regulator CtrA at the -35 position; this motif was footprinted by purified Caulobacter crescentus CtrA protein in its phosphorylated state. We have succeeded in isolating synchronized populations of Agrobacterium cells and analyzing their progression through the cell cycle. We demonstrate that DNA replication and cell division can be followed in an orderly manner and that flagellin expression is cyclic, consistent with our observation that motility varies during the cell cycle. Using these synchronized populations, we show that CcrM methylation of the chromosome is restricted to the late S phase of the cell cycle. Thus, within the alpha subdivision, there is a conserved cell cycle dependence and regulatory mechanism controlling ccrM expression.
Authors:
L S Kahng; L Shapiro
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Publication Detail:
Type:  Journal Article; Research Support, U.S. Gov't, Non-P.H.S.; Research Support, U.S. Gov't, P.H.S.    
Journal Detail:
Title:  Journal of bacteriology     Volume:  183     ISSN:  0021-9193     ISO Abbreviation:  J. Bacteriol.     Publication Date:  2001 May 
Date Detail:
Created Date:  2001-04-30     Completed Date:  2001-05-31     Revised Date:  2009-11-18    
Medline Journal Info:
Nlm Unique ID:  2985120R     Medline TA:  J Bacteriol     Country:  United States    
Other Details:
Languages:  eng     Pagination:  3065-75     Citation Subset:  IM    
Affiliation:
Department of Developmental Biology, Stanford University School of Medicine, Stanford, CA 94305, USA.
Data Bank Information
Bank Name/Acc. No.:
GENBANK/AF327563
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MeSH Terms
Descriptor/Qualifier:
Amino Acid Sequence
Base Sequence
Cell Cycle / physiology*
Culture Media
DNA Methylation*
Flow Cytometry
Gene Expression Regulation, Bacterial*
Molecular Sequence Data
Promoter Regions, Genetic
Rhizobium radiobacter / enzymology*,  growth & development,  physiology
Site-Specific DNA-Methyltransferase (Adenine-Specific) / chemistry,  genetics*,  metabolism
Grant Support
ID/Acronym/Agency:
GM51426/GM/NIGMS NIH HHS; K08-AI-01510/AI/NIAID NIH HHS
Chemical
Reg. No./Substance:
0/Culture Media; EC 2.1.1.-/DNA modification methylase CcrM; EC 2.1.1.72/Site-Specific DNA-Methyltransferase (Adenine-Specific)
Comments/Corrections

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