Document Detail


Bioinformatic and biochemical studies point to AAGR-1 as the ortholog of human acid alpha-glucosidase in Caenorhabditis elegans.
MedLine Citation:
PMID:  20349118     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Human acid alpha-glucosidase (GAA, EC 3.2.1.20) is a lysosomal enzyme that belongs to the glycoside hydrolase family 31 (GH31) and catalyses the hydrolysis of alpha-1,4- and alpha-1,6-glucosidic linkages at acid pH. Hereditary deficiency of GAA results in lysosomal glycogen storage disease type II (GSDII, Pompe disease). The aim of this study was to assess GH31 proteins in Caenorhabditis elegans (C. elegans) to identify the ortholog of human GAA. Bioinformatic searches for GAA ortholog in C. elegans genome revealed four acid alpha-glucosidase-related (aagr-1-4) genes. Multiple sequence alignment of AAGRs with other GH31 proteins demonstrated their evolutionary conservation. Phylogenetic analyses suggested clustering of AAGR-1 and -2 with acid-active and AAGR-3 and -4 with neutral-active GH31 enzymes. In order to prove the AAGRs' predicted alpha-glucosidase activity, we performed RNA interference of all four aagr genes. The impact on the alpha-glucosidase activity was evaluated at pH 4.0 (acid) and pH 6.5 (neutral), with or without the inhibitor acarbose. AAGR-1 and -2 expressed acidic alpha-glucosidase activity; on the contrary, AAGR-3 not -4 represented the predominant neutral alpha-glucosidase activity in C. elegans. Similar results were obtained in each of aagr-1 and -4 deletion mutants. Moreover, based on our structural models of AAGRs and these biochemical experiments, we hypothesize that the enzymatic sensitivity of AAGR-2 and human maltase-glucoamylase to the inhibitor acarbose is associated with a tyrosine residue in the GH31 active site, whereas acarbose resistance of AAGR-1 and human GAA is associated with the corresponding tryptophane in the active site. Acid-active AAGR-1 may thus represent the ortholog of human GAA in C. elegans.
Authors:
Jakub Sikora; Jana Urinovská; Filip Majer; Helena Poupetová; Jitka Hlavatá; Marta Kostrouchová; Jana Ledvinová; Martin Hrebícek
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2010-03-27
Journal Detail:
Title:  Molecular and cellular biochemistry     Volume:  341     ISSN:  1573-4919     ISO Abbreviation:  Mol. Cell. Biochem.     Publication Date:  2010 Aug 
Date Detail:
Created Date:  2010-07-26     Completed Date:  2010-12-06     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  0364456     Medline TA:  Mol Cell Biochem     Country:  Netherlands    
Other Details:
Languages:  eng     Pagination:  51-63     Citation Subset:  IM    
Affiliation:
1st Faculty of Medicine, Institute of Inherited Metabolic Disorders, Charles University in Prague, Ke Karlovu 2, Prague 2, Czech Republic. jakub.sikora@lf1.cuni.cz
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MeSH Terms
Descriptor/Qualifier:
Acarbose / pharmacology
Animals
Caenorhabditis elegans Proteins / antagonists & inhibitors,  chemistry,  genetics*
Catalytic Domain
Computational Biology / methods
Humans
Phylogeny
Sequence Alignment
alpha-Glucosidases / antagonists & inhibitors,  chemistry,  genetics*
Chemical
Reg. No./Substance:
0/Caenorhabditis elegans Proteins; 56180-94-0/Acarbose; EC 3.2.1.20/Aagr-1 protein, C elegans; EC 3.2.1.20/GAA protein, human; EC 3.2.1.20/alpha-Glucosidases

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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