Document Detail


Association of ACAT1-positive vesicles with late endosomes/ lysosomes in cholesterol-rich human macrophages.
MedLine Citation:
PMID:  20523008     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
AIM: Acyl-coenzyme A: cholesterol acyltransferase1 (ACAT1) is an endoplasmic reticulum (ER)-resident enzyme that catalyzes the conversion of cholesterol into cholesteryl esters. We previously showed that cholesterol-loaded macrophages produce numerous ER-derived vesicles with elevated ACAT1 enzyme activity; some of these vesicles were shown to be closely associated with Golgi-related organelle(s). The aim of this study was to investigate the translocation of ACAT1 vesicle in cholesterol-loaded macrophages.
METHODS: To demonstrate association of ACAT1 with late endosomes/lysosomes (LE/LS), primary human macrophages with or without cholesterol-loading was subjected to confocal microscopy, immunoelectron microscopy, subcellular fractionation, and immunoadsorption assay. Furthermore, cholesterol esterification assay was also carried out to investigate function of ACAT1 associated LE/LS.
RESULTS: Confocal fluorescence microscopy revealed that no significant ACAT1 signal was associated with the signal for LAMP2, a marker protein for LE/LS, in cholesterol non-loaded macrophages; however, approximately 20% of the total ACAT1 signals colocalized with the LAMP2 signal in cholesterol-loaded macrophages. ACAT1-positive membranes isolated by immunoadsorption using ACAT1-specific antibody contained LAMP2, demonstrating the association of ACAT1 and LE/LS. In addition, in macrophages phagocytosing latex beads, the close association of ACAT1 with LE/LS can be demonstrated in phagosomes isolated from cholesterol-loaded macrophages, not from non-loaded macrophages. Furthermore, cholesterol-loaded macrophages re-esterified aggregated LDL-derived cholesteryl ester even in the presence of U18666A, a reagent known to block egression of cholesterol from LE/LS.
CONCLUSION: Our results indicated that cholesterol-loaded human macrophages produce LE/LS in close association with ACAT1, and may promote efficient esterification of modified LDL-derived free cholesterol on LE/LS.
Authors:
XiaoFeng Lei; Yukio Fujiwara; Catherine C Y Chang; Ta-Yuan Chang; Motohiro Takeya; Naomi Sakashita
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2010-06-02
Journal Detail:
Title:  Journal of atherosclerosis and thrombosis     Volume:  17     ISSN:  1880-3873     ISO Abbreviation:  J. Atheroscler. Thromb.     Publication Date:  2010 Jul 
Date Detail:
Created Date:  2010-08-04     Completed Date:  2010-11-16     Revised Date:  2011-07-01    
Medline Journal Info:
Nlm Unique ID:  9506298     Medline TA:  J Atheroscler Thromb     Country:  Japan    
Other Details:
Languages:  eng     Pagination:  740-50     Citation Subset:  IM    
Affiliation:
Department of Cell Pathology, Graduate School of Medical Sciences, Kumamoto University, 1-1-1 Honjo, Kumamoto, Japan.
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MeSH Terms
Descriptor/Qualifier:
Acetyl-CoA C-Acetyltransferase / metabolism*
Animals
Cells, Cultured
Cholesterol / metabolism*
Endoplasmic Reticulum / metabolism*
Endosomes / metabolism*
Esterification
Humans
Immunoblotting
Lysosomal-Associated Membrane Protein 2 / metabolism
Lysosomes / metabolism*
Macrophages / metabolism*
Mice
Subcellular Fractions
Chemical
Reg. No./Substance:
0/Lysosomal-Associated Membrane Protein 2; 57-88-5/Cholesterol; EC 2.3.1.9/ACAT1 protein, human; EC 2.3.1.9/Acat1 protein, mouse; EC 2.3.1.9/Acetyl-CoA C-Acetyltransferase

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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