Document Detail


Aryl hydrocarbon receptor nuclear translocator/hypoxia-inducible factor-1{beta} plays a critical role in maintaining glucose-stimulated anaplerosis and insulin release from pancreatic {beta}-cells.
MedLine Citation:
PMID:  21059654     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
The metabolic pathways that are involved in regulating insulin secretion from pancreatic β-cells are still incompletely understood. One potential regulator of the metabolic phenotype of β-cells is the transcription factor aryl hydrocarbon receptor nuclear translocator (ARNT)/hypoxia-inducible factor (HIF)-1β. ARNT/HIF-1β levels are profoundly reduced in islets obtained from type 2 diabetic patients. However, no study to date has investigated key pathways involved in regulating insulin release in β-cells that lack ARNT/HIF-1β. In this study, we confirm that siRNA-mediated knockdown of ARNT/HIF-1β inhibits glucose-stimulated insulin secretion. We next investigated the metabolic consequence of the loss of ARNT/HIF-1β knockdown. We demonstrate that β-cells with reduced ARNT/HIF-1β expression levels exhibit a 31% reduction in glycolytic flux without significant changes in glucose oxidation or the ATP:ADP ratio. Metabolic profiling of β-cells treated with siRNAs against the ARNT/HIF-1β gene revealed that glycolysis, anaplerosis, and glucose-induced fatty acid production were down-regulated, and all are key events involved in glucose-stimulated insulin secretion. In addition, both first and second phase insulin secretion in islets were significantly reduced after ARNT/HIF-1β knockdown. Together, our data suggest an important role for ARNT/HIF-1β in anaplerosis, and it may play a critical role in maintaining normal secretion competence of β-cells.
Authors:
Renjitha Pillai; Peter Huypens; Mei Huang; Stephanie Schaefer; Tanya Sheinin; Shawn D Wettig; Jamie W Joseph
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2010-11-08
Journal Detail:
Title:  The Journal of biological chemistry     Volume:  286     ISSN:  1083-351X     ISO Abbreviation:  J. Biol. Chem.     Publication Date:  2011 Jan 
Date Detail:
Created Date:  2011-01-10     Completed Date:  2011-03-02     Revised Date:  2012-01-16    
Medline Journal Info:
Nlm Unique ID:  2985121R     Medline TA:  J Biol Chem     Country:  United States    
Other Details:
Languages:  eng     Pagination:  1014-24     Citation Subset:  IM    
Affiliation:
School of Pharmacy, University of Waterloo, Waterloo, Ontario N2L 3G1, Canada.
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MeSH Terms
Descriptor/Qualifier:
Adenosine Diphosphate / metabolism
Adenosine Triphosphate / metabolism
Animals
Aryl Hydrocarbon Receptor Nuclear Translocator / genetics,  metabolism*
Cell Line, Tumor
Citric Acid Cycle / physiology
Diabetes Mellitus, Type 2 / genetics,  metabolism*
Fatty Acids, Nonesterified / metabolism
Gas Chromatography-Mass Spectrometry
Gene Expression Profiling
Glucose / metabolism*,  pharmacology
Insulin / metabolism*
Insulin-Secreting Cells / cytology,  metabolism*
Insulinoma
Metabolomics
Oxidation-Reduction
Pancreatic Neoplasms
Pentose Phosphate Pathway / physiology
RNA, Small Interfering
Rats
Grant Support
ID/Acronym/Agency:
//Canadian Institutes of Health Research
Chemical
Reg. No./Substance:
0/ARNT protein, rat; 0/Fatty Acids, Nonesterified; 0/Insulin; 0/RNA, Small Interfering; 138391-32-9/Aryl Hydrocarbon Receptor Nuclear Translocator; 50-99-7/Glucose; 56-65-5/Adenosine Triphosphate; 58-64-0/Adenosine Diphosphate

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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