Document Detail


Aryl hydrocarbon receptor-mediated induction of Stearoyl-CoA desaturase 1 alters hepatic fatty acid composition in TCDD-elicited steatosis.
MedLine Citation:
PMID:  21890736     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
2,3,7,8-tetrachlorodibenzo-ρ-dioxin (TCDD) induces hepatic dyslipidemia mediated by the aryl hydrocarbon receptor (AhR). Stearoyl-CoA desaturase 1 (Scd1) performs the rate-limiting step in monounsaturated fatty acid (MUFA) synthesis, desaturating 16:0 and 18:0 into 16:1n7 and 18:1n9, respectively. To further examine the role of Scd1 in TCDD-induced hepatotoxicity, comparative studies were performed in Scd1(+/+) and Scd1(-/-) mice treated with 30 μg/kg TCDD. TCDD induced Scd1 activity, protein, and messenger RNA (mRNA) levels approximately twofold. In Scd1(+/+) mice, hepatic effects were marked by increased vacuolization and inflammation and a 3.5-fold increase in serum alanine aminotransferase (ALT) levels. Hepatic triglycerides (TRGs) were induced 3.9-fold and lipid profiling by gas chromatography-mass spectroscopy measured a 1.9-fold increase in fatty acid (FA) levels, consistent with the induction of lipid transport genes. Induction of Scd1 altered FA composition by decreasing saturated fatty acid (SFA) molar ratios 8% and increasing MUFA molar ratios 9%. Furthermore, ChIP-chip analysis revealed AhR enrichment (up to 5.7-fold), and computational analysis identified 16 putative functional dioxin response elements (DREs) within Scd1 genomic loci. Band shift assays confirmed AhR binding with select DREs. In Scd1(-/-) mice, TCDD induced minimal hepatic vacuolization and inflammation, while serum ALT levels remained unchanged. Although Scd1 deficiency attenuated TCDD-induced TRG accumulation, overall FA levels remained unchanged compared with Scd1(+/+) mice. In Scd1(-/-) mice, TCDD induced SFA ratios 8%, reduced MUFA ratios 13%, and induced polyunsaturated fatty acid ratios 5% relative to treated Scd1(+/+) mice. Collectively, these results suggest that AhR regulation of Scd1 not only alters lipid composition but also contributes to the hepatotoxicity of TCDD.
Authors:
Michelle M Angrish; A D Jones; Jack R Harkema; Timothy R Zacharewski
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural     Date:  2011-09-02
Journal Detail:
Title:  Toxicological sciences : an official journal of the Society of Toxicology     Volume:  124     ISSN:  1096-0929     ISO Abbreviation:  Toxicol. Sci.     Publication Date:  2011 Dec 
Date Detail:
Created Date:  2011-11-17     Completed Date:  2012-05-16     Revised Date:  2012-05-24    
Medline Journal Info:
Nlm Unique ID:  9805461     Medline TA:  Toxicol Sci     Country:  United States    
Other Details:
Languages:  eng     Pagination:  299-310     Citation Subset:  IM    
Affiliation:
Genetics Program, Michigan State University, East Lansing, Michigan 48824, USA.
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MeSH Terms
Descriptor/Qualifier:
Animals
Blotting, Western
Drug-Induced Liver Injury / genetics,  metabolism
Fatty Acids / metabolism*
Fatty Liver / genetics,  metabolism*
Gas Chromatography-Mass Spectrometry
Liver / drug effects*,  metabolism
Mice
Mice, Knockout
Principal Component Analysis
Real-Time Polymerase Chain Reaction
Receptors, Aryl Hydrocarbon / metabolism*
Response Elements / genetics
Stearoyl-CoA Desaturase / biosynthesis*,  genetics
Tetrachlorodibenzodioxin / toxicity*
Triglycerides / metabolism
Grant Support
ID/Acronym/Agency:
P42ES04911/ES/NIEHS NIH HHS
Chemical
Reg. No./Substance:
0/Fatty Acids; 0/Receptors, Aryl Hydrocarbon; 0/Triglycerides; 1746-01-6/Tetrachlorodibenzodioxin; EC 1.14.19.1/Scd1 protein, mouse; EC 1.14.19.1/Stearoyl-CoA Desaturase

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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