Document Detail


Anthrax toxin induces macrophage death by p38 MAPK inhibition but leads to inflammasome activation via ATP leakage.
MedLine Citation:
PMID:  21683629     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Detection of microbial constituents by membrane associated and cytoplasmic pattern recognition receptors is the essence of innate immunity, leading to activation of protective host responses. However, it is still unclear how immune cells specifically respond to pathogenic bacteria. Using virulent and nonvirulent strains of Bacillus anthracis, we have shown that secretion of ATP by infected macrophages and the sequential activation of the P2X7 purinergic receptor and nucleotide binding oligomerization domain (NOD)-like receptors are critical for IL-1-dependent host protection from virulent B. anthracis. Importantly, lethal toxin produced by virulent B. anthracis blocked activation of protein kinases, p38 MAPK and AKT, resulting in opening of a connexin ATP release channel and induction of macrophage death. Prevention of cell death or ATP release through constitutive p38 or AKT activation interfered with inflammasome activation and IL-1β production, thereby compromising antimicrobial immunity.
Authors:
Syed Raza Ali; Anjuli M Timmer; Sameera Bilgrami; Eek Joong Park; Lars Eckmann; Victor Nizet; Michael Karin
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't     Date:  2011-06-16
Journal Detail:
Title:  Immunity     Volume:  35     ISSN:  1097-4180     ISO Abbreviation:  Immunity     Publication Date:  2011 Jul 
Date Detail:
Created Date:  2011-07-22     Completed Date:  2011-10-04     Revised Date:  2012-02-29    
Medline Journal Info:
Nlm Unique ID:  9432918     Medline TA:  Immunity     Country:  United States    
Other Details:
Languages:  eng     Pagination:  34-44     Citation Subset:  IM    
Copyright Information:
Copyright © 2011 Elsevier Inc. All rights reserved.
Affiliation:
Laboratory of Signal Transduction, Department of Pharmacology, University of California, San Diego, La Jolla, California, USA.
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MeSH Terms
Descriptor/Qualifier:
Adaptor Proteins, Signal Transducing / metabolism
Adenosine Triphosphate / secretion
Animals
Anthrax / immunology*,  microbiology
Antigens, Bacterial / genetics,  immunology,  metabolism*
Apoptosis / drug effects
Apoptosis Regulatory Proteins / metabolism
Bacillus anthracis / genetics,  immunology*,  pathogenicity
Bacterial Toxins / genetics,  immunology,  metabolism*
Cells, Cultured
Connexin 43 / metabolism
Immunity, Innate / genetics
Inflammasomes / metabolism*
Interleukin-1beta / immunology,  metabolism
Macrophages, Peritoneal / immunology,  metabolism*,  microbiology,  pathology
Mice
Mice, Inbred C57BL
Mutation / genetics
Oncogene Protein v-akt / antagonists & inhibitors
Receptors, Purinergic P2X7 / metabolism
Virulence / genetics
p38 Mitogen-Activated Protein Kinases / antagonists & inhibitors
Grant Support
ID/Acronym/Agency:
R01 AI043477-15/AI/NIAID NIH HHS; R01 ES006376-19/ES/NIEHS NIH HHS
Chemical
Reg. No./Substance:
0/Adaptor Proteins, Signal Transducing; 0/Antigens, Bacterial; 0/Apoptosis Regulatory Proteins; 0/Bacterial Toxins; 0/Connexin 43; 0/Inflammasomes; 0/Interleukin-1beta; 0/NALP1 protein, mouse; 0/Receptors, Purinergic P2X7; 0/anthrax toxin; 56-65-5/Adenosine Triphosphate; EC 2.7.11.1/Oncogene Protein v-akt; EC 2.7.11.24/p38 Mitogen-Activated Protein Kinases

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