Document Detail


Activity-dependent NFATc3 nuclear accumulation in pericytes from cortical parenchymal microvessels.
MedLine Citation:
PMID:  17881610     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
The calcium-dependent transcription factor NFATc3, which is a member of the nuclear factor of activated T cells (NFAT) family of transcription factors, is critical for embryonic vascular development and differentiation. Despite its potential importance, nothing is known about NFATc3 regulation in the brain microcirculation. In the present study, we sought to investigate the role that glutamate, possibly through astrocytic communication, plays in the control of NFATc3 regulation in pericytes from parenchymal microvessels. Coronal cortical slices from neonatal rats were subjected to electrical field stimulation or were treated with the metabotropic glutamate receptor agonist (+/-)-1-aminocyclopentane-trans-1,3-dicarboxylic acid (t-ACPD). NFATc3, glial fibrillary acidic protein (an astrocyte-specific marker), and platelet-derived growth factor-beta-receptor (a pericyte-specific marker) were detected by immunofluorescence. Electrical field stimulation induced NFATc3 nuclear accumulation in pericytes. This response was dependent on neuronal activity and group I metabotropic glutamate receptor (mGluR) activation. In addition, t-ACPD significantly increased NFATc3 nuclear accumulation in both astrocytes and pericytes. NFATc3 nuclear accumulation in pericytes was prevented when astrocytic function was abolished with the gliotoxin L-alpha-aminoadipate or by the inhibition of calcineurin, cyclooxygenase, and nitric oxide synthase. This is the first study to report NFATc3 expression in pericytes from parenchymal microvessels and in astrocytes from native tissue. Our results suggest a model by which glutamate, via mGluR activation, may regulate gene transcription in pluripotent vascular pericytes.
Authors:
Jessica A Filosa; Mark T Nelson; Laura V Gonzalez Bosc
Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't     Date:  2007-09-19
Journal Detail:
Title:  American journal of physiology. Cell physiology     Volume:  293     ISSN:  0363-6143     ISO Abbreviation:  Am. J. Physiol., Cell Physiol.     Publication Date:  2007 Dec 
Date Detail:
Created Date:  2007-12-12     Completed Date:  2008-03-04     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  100901225     Medline TA:  Am J Physiol Cell Physiol     Country:  United States    
Other Details:
Languages:  eng     Pagination:  C1797-805     Citation Subset:  IM    
Affiliation:
Department of Psychiatry, University of Cincinnati, Cincinnati, OH 45237, USA. jessica.filosa@uc.edu
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MeSH Terms
Descriptor/Qualifier:
Animals
Animals, Newborn
Astrocytes / metabolism,  physiology*
Cell Nucleus / metabolism*
Cerebral Cortex / blood supply
Electric Stimulation
Gene Expression Regulation
Glutamic Acid / metabolism*
NFATC Transcription Factors / metabolism*
Pericytes / metabolism*
Rats
Rats, Sprague-Dawley
Receptors, Metabotropic Glutamate / agonists
Signal Transduction / physiology
Grant Support
ID/Acronym/Agency:
DK53832/DK/NIDDK NIH HHS; HL44455/HL/NHLBI NIH HHS; HL63722/HL/NHLBI NIH HHS
Chemical
Reg. No./Substance:
0/NFATC Transcription Factors; 0/Receptors, Metabotropic Glutamate; 0/transcription factor NF-AT c3; 56-86-0/Glutamic Acid

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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