Document Detail


Absence of both IL-7 and IL-15 severely impairs the development of CD8 T cell response against Toxoplasma gondii.
MedLine Citation:
PMID:  20520779     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
CD8(+) T cells play an essential role in the protection against both acute as well as chronic Toxoplasma gondii infection. Although the role of IL-15 has been reported to be important for the development of long-term CD8(+) T cell immunity against the pathogen, the simultaneous roles played by both IL-15 and related gamma-chain family cytokine IL-7 in the generation of this response during acute phase of infection has not been described. We demonstrate that while lack of IL-7 or IL-15 alone has minimal impact on splenic CD8(+) T cell maturation or effector function development during acute Toxoplasmosis, absence of both IL-7 and IL-15 only in the context of infection severely down-regulates the development of a potent CD8(+) T cell response. This impairment is characterized by reduction in CD44 expression, IFN-gamma production, proliferation and cytotoxicity. However, attenuated maturation and decreased effector functions in these mice are essentially downstream consequences of reduced number of antigen-specific CD8(+) T cells. Interestingly, the absence of both cytokines did not impair initial CD8(+) T cell generation but affected their survival and differentiation into memory phenotype IL-7Ralpha(hi) cells. Significantly lack of both cytokines severely affected expression of Bcl-2, an anti-apoptotic protein, but minimally affected proliferation. The overarching role played by these cytokines in eliciting a potent CD8(+) T cell immunity against T. gondii infection is further evidenced by poor survival and high parasite burden in anti IL-7 treated IL-15(-/-) mice. These studies demonstrate that the two cytokines, IL-7 and IL-15, are exclusively important for the development of protective CD8(+) T cell immune response against T. gondii. To the best of our knowledge this synergism between IL-7 and IL-15 in generating an optimal CD8(+) T cell immunity against intracellular parasite or any other infectious disease model has not been previously reported.
Authors:
Rajarshi Bhadra; Hongbing Guan; Imtiaz A Khan
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural     Date:  2010-05-26
Journal Detail:
Title:  PloS one     Volume:  5     ISSN:  1932-6203     ISO Abbreviation:  PLoS ONE     Publication Date:  2010  
Date Detail:
Created Date:  2010-06-03     Completed Date:  2010-09-07     Revised Date:  2010-09-28    
Medline Journal Info:
Nlm Unique ID:  101285081     Medline TA:  PLoS One     Country:  United States    
Other Details:
Languages:  eng     Pagination:  e10842     Citation Subset:  IM    
Affiliation:
Department of Microbiology, Immunology and Tropical Medicine, George Washington University, Washington, DC., USA.
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MeSH Terms
Descriptor/Qualifier:
Animals
Antibodies / pharmacology
Bromodeoxyuridine / metabolism
CD8-Positive T-Lymphocytes / cytology,  drug effects,  immunology*,  parasitology*
Cell Cycle / drug effects
Down-Regulation / drug effects
Interferon-gamma / immunology
Interleukin-15 / deficiency*,  metabolism
Interleukin-7 / deficiency*,  immunology
Mice
Mice, Knockout
Parasites / drug effects,  immunology
Proto-Oncogene Proteins c-bcl-2 / metabolism
Receptors, Interleukin-7 / metabolism
Spleen / drug effects,  immunology,  parasitology
Staining and Labeling
Toxoplasma / drug effects,  immunology*
Toxoplasmosis / immunology,  parasitology
Up-Regulation / drug effects
Vaccination
Grant Support
ID/Acronym/Agency:
AI33325/AI/NIAID NIH HHS
Chemical
Reg. No./Substance:
0/Antibodies; 0/Interleukin-15; 0/Interleukin-7; 0/Proto-Oncogene Proteins c-bcl-2; 0/Receptors, Interleukin-7; 59-14-3/Bromodeoxyuridine; 82115-62-6/Interferon-gamma
Comments/Corrections

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