Document Detail


AUF1 cell cycle variations define genomic DNA methylation by regulation of DNMT1 mRNA stability.
MedLine Citation:
PMID:  17030625     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
DNA methylation is a major determinant of epigenetic inheritance. DNA methyltransferase 1 (DNMT1) is the enzyme responsible for the maintenance of DNA methylation patterns during cell division, and deregulated expression of DNMT1 leads to cellular transformation. We show herein that AU-rich element/poly(U)-binding/degradation factor 1 (AUF1)/heterogeneous nuclear ribonucleoprotein D interacts with an AU-rich conserved element in the 3' untranslated region of the DNMT1 mRNA and targets it for destabilization by the exosome. AUF1 protein levels are regulated by the cell cycle by the proteasome, resulting in cell cycle-specific destabilization of DNMT1 mRNA. AUF1 knock down leads to increased DNMT1 expression and modifications of cell cycle kinetics, increased DNA methyltransferase activity, and genome hypermethylation. Concurrent AUF1 and DNMT1 knock down abolishes this effect, suggesting that the effects of AUF1 knock down on the cell cycle are mediated at least in part by DNMT1. In this study, we demonstrate a link between AUF1, the RNA degradation machinery, and maintenance of the epigenetic integrity of the cell.
Authors:
Jerome Torrisani; Alexander Unterberger; Sachin R Tendulkar; Keisuke Shikimi; Moshe Szyf
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2006-10-09
Journal Detail:
Title:  Molecular and cellular biology     Volume:  27     ISSN:  0270-7306     ISO Abbreviation:  Mol. Cell. Biol.     Publication Date:  2007 Jan 
Date Detail:
Created Date:  2006-12-14     Completed Date:  2007-01-18     Revised Date:  2009-11-18    
Medline Journal Info:
Nlm Unique ID:  8109087     Medline TA:  Mol Cell Biol     Country:  United States    
Other Details:
Languages:  eng     Pagination:  395-410     Citation Subset:  IM    
Affiliation:
Department of Pharmacology and Therapeutics, McGill University, 3655 Sir William Osler Promenade, Montreal, Quebec H3G 1Y6, Canada.
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MeSH Terms
Descriptor/Qualifier:
3' Untranslated Regions
Amino Acid Sequence
Animals
Base Sequence
Cell Cycle
DNA (Cytosine-5-)-Methyltransferase / metabolism,  physiology*
DNA Methylation*
Epigenesis, Genetic
Gene Expression Regulation*
Hela Cells
Heterogeneous-Nuclear Ribonucleoprotein D / metabolism,  physiology*
Humans
Mice
Mice, Inbred BALB C
Molecular Sequence Data
Chemical
Reg. No./Substance:
0/3' Untranslated Regions; 0/Heterogeneous-Nuclear Ribonucleoprotein D; 0/hnRNP D0; EC 2.1.1.37/DNA (Cytosine-5-)-Methyltransferase; EC 2.1.1.37/DNA (cytosine-5-)-methyltransferase 1
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