Revision of the genus Euscelidia Westwood, 1850 (Diptera: Asilidae: Leptogastrinae).

Revision of the genus Euscelidia Westwood, 1850 (Diptera: Asilidae: Leptogastrinae).

Article Type:	Report
Subject:	Asilidae (Identification and classification) Asilidae (Natural history)
Author:	Dikow, Torsten
Pub Date:	12/01/2003
Publication:	Name: African Invertebrates Publisher: The Council of Natal Museum Audience: Academic Format: Magazine/Journal Subject: Zoology and wildlife conservation Copyright: COPYRIGHT 2003 The Council of Natal Museum ISSN: 1681-5556
Issue:	Date: Dec, 2003 Source Volume: 44 Source Issue: 2
Geographic:	Geographic Scope: South Africa Geographic Code: 6SOUT South Africa

Accession Number:	208746198
Full Text:	Euscelidia cacula sp. n. Figs 17E-H, 46 Etymology: Noun in apposition that refers to the type locality Cacula, Angola. Diagnosis: The species is distinguished from congeners by the predominantly apruinose sct, many yellow setae arranged laterally on sct, and the densely arranged microtrichia on the wings. Description: Head: Black; fc yellow pruinose, fc gib indistinct, mystax yellow, many macrosetae; prob and plp brownish-black, setae white; oc tr yellow pruinose dorsally; occ silver pruinose, setae yellow, white setae ventrally; Antennae--scp dark brown, yellow setae ventrally, white pruinose; ped dark brown, brown setae ventrally and dorsally, white pruinose; pped brown, white pruinose proximally and ventrally; apsel hyaline. Thorax: Black; ppro peg large, yellow pruinose; sct apruinose, margins yellow pruinose, yellow setae on margins, few short dc setae may be present, macrosetae: 1 white npl s, 1 black spal s; sctl white pruinose, ds sctl s and sctl s long, yellow; Legs--yellow to brown; pro fern light brown, meso fern light brown, brown dorsally, meta fern yellow proximally, remaining part light brown to brown dorsally, light brown ventrally, slightly clubbed, short white setae on club; pro tib light brown, yellow stripe anteriorly, meso and meta tib brown, yellow stripe not reaching tip on meta tib anteriorly; first tar yellow proximally, brown distad, black setae; emp about a third of length of clw; Wings--densely covered with microtrichia; ptero indistinct, brown; cell d terminating in 2 veins; hlt orange. Abdomen: Black; T grey pruinose, distal T more extensively brown pruinose, T1 with long yellow setae, T2 in proximal 2/3 with yellow setae laterally, remaining T with short white and yellow setae, S grey pruinose; [male] terminalia Figs 17E-H--sur pointed distally, lobe ventrally; hypd ventral margin straight; d aed shea tubular, terminating in a dorsad directed structure; lat apod bifurcated. Type material--The [male] holotype is labelled 'ANGOLA (A20) 10 mls [16 km], NE. Cacula 5.iii.1972 ('ANGOLA (A20)' blue underlined) / Southern Africa Exp. B.M. 1972-1 / Euscelidia? sp. nov. det. H. Oldroyd 1973 (handwritten except for 'det. H. Oldroyd 19') / HOLOTYPE Euscelidia cacula sp. nov. T. Dikow 2001 (red label)'. The specimen is double mounted (minuten in block of foam), is in very good condition (right pped broken; [male] terminalia attached to specimens pin in micro vial), and is deposited in the BMNH. A paratype of undeterminable gender with the same locality data and 'PARATYPE Euscelidia cacula sp. nov. T. Dikow 2001 (yellow label)' is double mounted (minuten in block of foam), is in poor condition (antennae, left meta leg, wings, and abdomen broken), and is deposited in the BMNH. A [female] paratype is labelled 'ANGOLA (A 17) 15 mls [24 km]. N. Sa da Bandeira, c. 6500 ft. [1950 m] 3.iii.1972 ('ANGOLA (AI7)' blue underlined) / Southern Africa Exp. B.M. 1972-1 / PARATYPE Euscelidia cacula sp. nov. T. Dikow 2001 (yellow label)'. The specimen is double mounted (minuten in block of foam), is in very good condition (pro and meso tarsi broken), and is deposited in the BMNH. Another [female] paratype is labelled 'ANGOLA (A22) Nova Lisboa 5-7.iii.1972 ('ANGOLA (A22)' blue underlined) / Southern Africa Exp. B.M. 1972-1 / PARATYPE Euscelidia cacula sp. nov. T. Dikow 2001 (yellow label)'. The specimen is double mounted (minuten in block of foam), is in excellent condition, and is deposited in the BMNH. Type locality and distribution (Fig. 46): Angola, Cacula, 14[degrees]29'S 14[degrees]06'E. Angola. Euscelidia castanea Janssens, 1954 Fig. 50 Euscelidia castanea Janssens, 1954a: 124; Oldroyd 1980: 356. Diagnosis: The species is distinguished, together with nenemusha and schoutedeni, from congeners by the overall brown colour, the predominantly apruinose sct, and the apruinose T8 in the [female] [female]. Redescription: Head: Black; fc silver pruinose, fc gib indistinct, mystax white, 6 macrosetae; oc tr white pruinose dorsally; occ silver pruinose, few white setae; Antennae--scp and ped brown, setae brown; pped brown, white pruinose; apsel brown; Thorax: Brown; sct brown, predominantly apruinose, anterior, lateral, and posterior margins silver pruinose, few white setae on pruinose area, brown setae on posterior margin, macrosetae: black, 1 npl s, 1 spal s; sctl silver pruinose, ds sctl s and sctl long, brown; Legs--yellow and light brown; fern yellow proximally, brown distad; tib light brown, yellow stripe not reaching tip anteriorly; emp short; Wings--hyaline, brown stained; ptero distinct, brown; cell d terminating in 2 veins; hlt brown; Abdomen: Brown; T predominantly apruinose, lateral and posterior margins white pruinose, T1 with black setae, T2 with white long setae laterally, remaining T with short white setae, [female] T8 apruinose; [male] terminalia--[male] [male] specimens unknown. Type material--The [female] holotype is labelled 'HOLOTYPE (red label with black submarginal border) /TYPE (red label with red border around 'TYPE') / Congo belge: E N. U. [Parc National de l'Upemba] Lusinga (1.760 m.) 1-8-XII-1947 Mis. G. F. de Witte. 1123a / E. Janssens det., 1952 Euscelidia castanea Em. Janssens (handwritten except for 'E. Janssens det., 19')'. The specimen is double mounted (minuten in block of foam), is in good condition (met legs broken and fight Wing damaged), and is deposited in the MRAC. Type locality and distribution (Fig. 50): DR Congo, Upemba National Park, Lusinga, 8[degrees]55'S 27[degrees]11'E. DR Congo, Uganda. Material examined: DR CONGO: 1 [female] Virunga National Park, Mutsora, 0[degrees] 19'N 29[degrees]44'E, 13-18.ii.1953; 1 [female] Upemba National Park, Lusinga, 8[degrees]55'S 27[degrees]11'E, 1-8.xii.1947; UGANDA: 1 [female] Kilembe, 0[degrees] 12'N29[degrees]59'E, xii.1934-i.1935. Depository: BMNH, ISNB, MRAC. Remarks: The species is only known from a few female specimens. It is related to nenemusha and schoutedeni, which are also only known from females. It is necessary to examine male specimens from the type localities of all three species to discuss a probable synonymy. Euscelidia cobice sp. n. Figs 24A-D, 36 Etymology: The specific name 'cobice' is a noun in apposition that refers to the Copenhagen Biosystematics Centre (COBICE) to acknowledge the generous funding by the European Commission 'Improving Human Potential' programme 'Transnational Access to Major Research Infrastructures' that made my research at this institution possible. Diagnosis: The species is distinguished from congeners by the predominantly yellow pruinose sct, the densely arranged microtrichia on the wings, and features of the terminalia (sur with large ventral lobe, d aed shea tubular). Description: Head: Black; fc silver pruinose, fc gib distinct, mystax white, many macrosetae; prob and pip brownish-black, setae white; oc tr apruinose; occ silver and yellowish pruinose, setae yellow; Antennae--scp brown, brown setae ventrally; ped brown, brown setae ventrally and dorsally, white pruinose; pped brown, white pruinose; apsel brown. Thorax: Black; ppro peg small, distinct, silver pruinose; sct bluish-black, predominantly yellow pruinose, median stripe and 2 lateral spots may be distinctly apruinose, sometimes fused to form 1 large apruinose spot, yellow setae scattered on surface; macrosetae: black, 1 npl s, 1 spal s; sctl yellow pruinose, ds sctl s and sctl s long, yellow; Legs yellow to brown; pro fem yellow, meso fem yellow proximally, remaining part brown, meta fern in proximal 1/3 pale yellow, clubbed, club in proximal half brown, distal half yellow, setae white, tib yellow proximally, remaining part brown, pale yellow stripe not reaching tip on meta tib anteriorly; first tar in proximal half yellow, brown in distal hall', remaining tar brown, black setae; emp short, about a quarter of length of clw; Wings--hyaline, but microtrichia evenly scattered on remigium, brown stained; ptero distinct, brown; cell d terminating in 2 veins; bit light brown. Abdomen: Brown, sometimes anterior margins of T orange; T predominantly brown pruinose, anterior, lateral, and posterior margins grey pruinose, T2 in proximal 1/3 with apruinose spot, T1 with long white and black setae laterally, T2 in proximal half with white setae laterally, remaining T with white and brown setae, S grey pruinose; [male] terminalia Figs 24A-D--sur pointed distally, large lobe ventrally; hypd ventral margin straight; d aed shea tubular, medium length; lat apod bifurcated. Type material--The [male] holotype is labelled 'S. India: Karnataka. Mudigere area, c. 900 m 2-10.xi.1977 Zool. Mus. Copenhagen Exp. / HOLOTYPE Euscelidia cobice sp. nov. det. T. Dikow 2001 (red label)'. The specimen is directly mounted, is in very good condition (right meta leg broken), and is deposited in the ZMUC. 7 [female] [female] 1 [male] paratypes are labelled 'S. India: Karnataka. Mudigere area, c. 900 m 2-10.xi.1977 Zool. Mus. Copenhagen Exp. / PARATYPE Euscelidia cobice sp. nov. det. T. Dikow 2001 (yellow label)'. The specimens are directly mounted, are in very good to excellent condition, and are deposited in the ZMUC. Type locality and distribution (Fig. 36): India, Karnataka Province, Mudigere, 13[degrees]07'N 75[degrees]37%. India. Material examined: INDIA: 2 [female] [female] Kodagu, 12[degrees]25'N 75[degrees]45'E, 24.x.-16.xi.1915. Depository: BMNH. [FIGURE 24 OMITTED] Euscelidia crena sp. n. Figs 24E-I, 29 Etymology: Latin noun crena = rounded projection; refers to the well-developed facial swelling. Diagnosis: The species is distinguished from congeners by the distinct facial swelling, three apruinose spots on the lateral sct, and features of the [male] terminalia (gonst large, lat apod triangular, d aed shea tubular). Description: Head: Black; fc silver pruinose, fc gib distinct, mystax white, 12 macrosetae; prob and pip brownish-black, setae white; oc tr white pruinose dorsally; occ silver pruinose, setae white; Antennae--scp brown, white setae ventrally; ped brown, brown setae ventrally and dorsally, white pruinose; pped light brown proximally, brown distad, white pruinose; apsel hyaline. Thorax: Dark brown; ppro peg small, distinct, silver pruinose; sct black ([female] paratype with orange anterior and lateral margins), yellow pruinose medially, margins silver pruinose, 2 antero-lateral spots apruinose, 2 lateral spots anterior and posterior to trn sut apruinose, but not as distinct as antero-lateral spots, few yellow or white setae on margins; macrosetae: white, 1 npl s, 1 spal s; sctl silver pruinose, ds sctl s and sctl s short, white; Legs--yellow to brown; pro and meso fern yellow, meta fern pale yellow in proximal half, clubbed, club light brown, setae white; pro and meso tib light brown, pale yellow stripe anteriorly, meta tib brown, pale yellow stripe in proximal half anteriorly; tar light brown to brown, black setae; emp long, about a quarter of length of clw; Wings--hyaline, few microtrichia scattered on remigium and distal margin; ptero distinct, light brown; cell d terminating in 3 veins; hlt light brown. Abdomen: Dark brown; T predominantly brown pruinose, anterior, lateral, and posterior margins grey pruinose, T1 with long white setae laterally, T2 in proximal third with few white setae laterally, remaining T with white and yellow setae, S grey pruinose; [male] terminalia Figs 24E-I--sur pointed distally, lobe ventrally; hypd ventral margin straight; gonst large; d aed shea tubular; aed apod small; lat apod triangular. Type material--The [male] holotype is labelled 'KENYA: Machakos #88 17 km SE Sultan Hamud 02[degrees]04'S:37[degrees]28'E 1140 m Date: 30.xi.1992 J Londt & A Whittington Grassland & road verges / HOLOTYPE Euscelidia crena sp. nov. det. T. Dikow 2001 (red label)'. The specimen is double mounted (minuten in block of foam), is in excellent condition ([male] terminalia attached to specimens pin in micro vial), and is deposited in the NMSA. The [female] paratype has same labels as holotype except 'PARATYPE Euscelidia crena sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is double mounted (minuten in block of foam), is in excellent condition, and is deposited in the NMSA. Type locality and distribution (Fig. 29): Kenya, Sultan Hamud (17 km SE), 2[degrees]04'S 37[degrees]28'E. Kenya. Euscelidia datis species-group Diagnosis of species: All representatives (artaphernes, datis, rapax, and two new species (lata, milva)) of this group are distinguished from congeners by the generally large size, darkened wings, the presence of yellow setae on the sct, erect setae on the clubbed meta fern (Fig. 25), and characters of the U terminalia (greatly enlarged, sharply inwards pointing tip of sur, very long aed (extending c; terminalia), and bifurcated lat apod). (Note: The use of this species-group does not intend any ranking of these species as a subgenus or even a genus.) Remarks: Only [male] [male] specimens of this group can be identified properly. The many [female] [female] specimens examined are listed under datis. The development of setae ventrad of the antennae in artaphernes is generally good for identifying [female] [female] of this species. However, the character is not consistently found in all [male] [male] specimens and therefore should be used tentatively. Some species of this group are sometimes found sympatrically, e.g. artaphernes and datis in Albertville (DR Congo) and Bossoum (Cameroon) and datis and milva in Fort Crampel (Central African Republic). It is necessary to examine more [male] [male] specimens to comment on the distribution of every species more accurately (see distribution map in Fig. 26). [FIGURE 25 OMITTED] Euscelidia artaphernes (Speiser, 1910) (E. datis species-group) Figs 26, 27A-C, G Leptogaster artaphernes Speiser, 1910: 83. Euscelidia artaphernes, Oldroyd 1939: 28; Lindner 1955: 28; Oldroyd 1980: 356. Redescription: Head: Black; fc yellow pruinose, often long yellow setae originating at antennal base directing ventrally, fc gib distinct; mystax yellow, many macrosetae, nearly as long as prob; oc tr apruinose; occ silver pruinose, setae yellow; Antennae--scp brown, setae yellow; ped light brown, white pruinose, setae yellow and brown; pped light brown proximally, brown distad, white pruinose posteriorly; apsel hyaline; Thorax: Black; sct predominantly black, antero-laterally orange; margins silver or yellow pruinose, yellow setae scattered on surface, setae always directing to the centre; sctl white pruinose, ds sctl s and sctl s long, yellow; Legs--orange to brown; fern light brown, meta fem pale yellow proximally, brown medially, orange distally, clubbed, club densely covered with short, erect yellow setae; pro and meso tib light brown, yellow stripe anteriorly, meta tib orange proximally, brown distad, yellow stripe not reaching tip anteriorly; emp distinct, very short; Wings--densely covered with microtrichia, brown coloured; ptero indistinct, brown; cell d terminating in 2 veins; hlt light brown; Abdomen: Brown to dark brown; T orange on anterior and posterior margins, grey pruinose, setae yellow, T1 with long yellow setae, T2 in proximal 2/3 with yellow setae laterally; [male] terminalia Figs 27A--C, G-sur bent inwards distally; hypd with membranous area ventrally; d aed shea long, tip Fig. 27G, lat apod bifurcated. Type material--The [male] lectotype, here designated to preserve stability and make more universal the use of this name, is labelled 'Kilimandjaro Sjostedt. 1905-6 / Kibonoto 1000-1300 m/29 april ('29' handwritten)/Type! Leptogaster artaphernes m P. Speiser det. (handwritten except for 'P. Speiser det.') / LECTOTYPE Leptogaster artaphernes Speiser, 1910 by T. Dikow 2001 (red label)'. The specimen is directly mounted, is in good condition (sct cracked), and is deposited in the NHRS. [FIGURE 26 OMITTED] [FIGURE 27 OMITTED] The [female] paralectotype is labelled 'Kilimandjaro Sjostedt. 1905-6 / Kibonoto 1000-1300 m / 4 dec. [4 December] ('4' handwritten) / Leptogaster artaphernes m P. Speiser det. (handwritten except for 'P. Speiser det.') / PARALECTOTYPE Leptogaster artaphernes Speiser, 1910 by T. Dikow 2001 (yellow label)'. The specimen is directly mounted, is in poor condition (right pped, left meso leg, right meta leg broken; sct cracked; left meta leg attached to the body with glue), and is deposited in the NHRS. Another paralectotype is labelled 'Kilimandjaro Sjostedt. 1905-6 / Obstgarten--Steppe / 22 Mars [March] ('22' handwritten)/Leptogaster artaphernes m P. Speiser det. (handwritten except for 'P. Speiser det.') / Paratypus (red label with black submarginal border) / 10 52 (red label) / PARALECTOTYPE Leptogaster artaphernes Speiser, 1910 by T. Dikow 2001 (yellow label)'. The specimen is directly mounted, is in poor condition (left pped, left meso leg, and abdomen broken (attached on a piece of cardboard); sct cracked), and is deposited in the NHRS. Type locality and distribution (Fig. 26): Tanzania, Kilimandjaro area, Kibonoto (= Kibongoto), 3[degrees]10'S 37[degrees]05'E. Burundi, Cameroon, DR Congo, Ethiopia, Kenya, Malawi, Mozambique, South Africa, Tanzania, Togo, Uganda, Zambia, Zimbabwe. Material examined: BURUNDI: 3 [male][male] Rumonge, 3[degrees]58'S 29[degrees]26'E, 21.i.1950; CAMEROON: 3 [male] [male] Bossoum, 9[degrees]56'N 13[degrees]41'E, 1.iv., 18.v., 1-10.vi.1914; DR CONGO: 1 [male] Kivu, 3[degrees]03'S 28[degrees]06'E; 2 [female] [female] Lulua, 5[degrees]55'S 25[degrees]46%, iv.1933; 1 [male] Albertville, 5[degrees]55'S 29[degrees]12%, xii.1918; 1 [male] Baudouinville, 7[degrees]03'S 29[degrees]47'E, 19.i.1933; ETHIOPIA: 2 [male] [male] Harer, 09[degrees]19'N 42[degrees]07'E; KENYA: 1 [male] Lake Victoria, 0[degrees]09'S 34[degrees]51'E, v.1912; 1 [female] Kuja Valley, 0[degrees]48'S 34[degrees]34'E, 30.iv.-1.v.1911; 2 [female] [female] Gelegele River, 0[degrees]53'S 35[degrees]02%, vi.1913; 1 [male] Nairobi, 1 [degrees] 16'S 36[degrees]48%, xi.-xii.1911; 1 [female] Kibwezi, 2[degrees]24'S 37[degrees]57'E, 2M-.iv.1911; MALAWI: 1 [female] Mponela, 13[degrees]30'S 33[degrees]42'E, 26.ii.1987; 1 [female] Senga Bay, 13[degrees]42'S 34[degrees]37'E, 7-8.iii.1987; 3 [female] [female] 1 [male] 1? Zomba, 15[degrees] 18'S 35[degrees] 10'E; MOZAMBIQUE: 1 [male] Nampula, 15[degrees]00'S 39[degrees]00'E, 19.ii.1982; SOUTH AFRICA: Northern Prov: 1 [male] Ofcolaco, 2430AB, 20.ii.1980; TANZANIA: 1 [female] Ukerewe, 2[degrees]01'S 32[degrees]58'E, 1911; 1 [female] Dar-es-Salam, 6[degrees]48'S 39[degrees]16'E, 11-20.xii.1961; 1 [male] vii.1902; 1 [male] NW Tanganyika, 1910; TOGO: 1 [male] Bismarckburg, 8[degrees]11'N 0[degrees]41'E, 9-11.v.1893; UGANDA: 1 [female] 2 [male] [male] Abi Farm, 3[degrees]05'N 30[degrees]56'E, 12.xi.1958; 1 [female] 2 [male] [male] Ankole, 0[degrees]43'N 30[degrees]11'E, 16.xii.1962, 23-29.iv.1968; 1 [female] 2 [male] [male] Mubende, 0[degrees]35'N 31 [degrees]22'E, 19+22.iv.1966; 1 [female] L. Namugabo, 0[degrees]04'N 32[degrees] 15'E, 3.vi.1963; 2 [female] [female] Bugalla Island, 0[degrees]38'S 32[degrees] 18'E, v.1912; 1 [female] 1 [male] 1? Entebbe, 0[degrees]03'N 32[degrees]27'E, 28.x.1912, 23.v.1913; 2 [male] [male] 1? Kome island, 0[degrees]05'S 32[degrees]44'E, 15.xii.1918; 2 [female] [female] Mbarara, 0[degrees]36'S 30[degrees]38'E, 22-24.x.1911, 15.xi.1934; 1 [female] 1 [male] Lolui Island, 0[degrees]07'S 33[degrees]42'E, v.1965; ZAMBIA: 1 [female] 1 [male] Kasanka National Game Reserve, 12[degrees]30'S 30[degrees]15'E, 16-22.xii.1989; 1 [female] Mungu, 15[degrees]16'S 23[degrees]04%, 10.xii.1955; ZIMBABWE: 1 [male] Harare, 17[degrees]46'S 31[degrees]02'E, i.1969; 1 [male] Bindura, 17[degrees]18'S 31[degrees]19'E, i.1929; COUNTRY UNKNOWN: 1 [female] Kawanda [Malawi, Uganda, or Zambia], 9.xi.1957; 1 [male] Abyssinia, xi.1911; 1 [male] no locality data. Depository: BMNH, ISNB, MRAC, NHMW, NMSA, SMNS, USNM, ZMHB, ZSMC. Remarks: Illustration of [male] terminalia by Theodor (1976: 20) is not artaphernes but datis. Euscelidia datis (Walker, 1849) (E. datis species-group) Figs 26, 28G-I, N Leptogaster datis Walker, 1849: 483; Hull 1962: 300. Euscelidia datis, Hermann 1926: 147; Oldroyd 1939: 28; Janssens 1953:6 1954b: 400: Theodor 1976: 20; Oldroyd 1980: 356. Lasiocnemuspilipes Curran, 1927: 2; Hull 1962: 309. syn. n. Euscelidia pilipes, Oldroyd 1980:357 (as junior synonym of rapax). Redescription: Head: Black; fc yellow pruinose, fc gib indistinct, mystax yellow, many macrosetae; oc tr apruinose; occ silver pruinose, setae yellow; Antennae -scp brown, setae yellow; ped light brown, setae yellow and brown, white pruinose; pped light brown proximally, brown distad, white pruinose; apsel hyaline; Thorax: Black; sct black, sometimes with orange-red spots antero-laterally, predominantly apruinose, margins white pruinose, yellow setae scattered on surface, macrosetae: white or black, 1 npl s, 1 spal s; sctl white pruinose, ds sctl s and sctl long, yellow; Legs--yellow to brown; fern light brown, meta fern pale yellow proximally, brown medially, clubbed, club densely covered with long, erect yellow setae; pro and meso tib light brown, yellow stripe anteriorly, meta tib in proximal half light brown, distal half brown, yellow stripe not reaching tip anteriorly; emp distinct, very short; Wings--densely covered with microtrichia, brown coloured; ptero indistinct, brown; cell d terminating in 2 veins; hlt yellow; Abdomen: Brown; T yellow pruinose, T1 with long yellow setae, T2 in proximal half with long yellow setae laterally; [male] terminalia Figs 28G-I, N (see also Figs 16-17 in Theodor (1976: 20))--sur bent inwards distally; hypd with membranous area distally; d aed shea long, tip simple, pointed (Fig. 28N), lat apod bifurcated. [FIGURE 28 OMITTED] Type material--The lectotype of Leptogaster datis of undeterminable gender, here designated to preserve stability and make more universal the use of this name, is labelled 'Type (circular label with green submarginal border) / One of Walker's series so named. Edw ('Edw' handwritten) Leptogaster datis Walk. (handwritten on backside) / Sierra Leone (handwritten) Presented by the Rev D.F. Morgan (on backside) / HOLOTYPE Leptogaster datis Walker det. J.E. CHAINEY 1982 (handwritten) Holotype (circular label with red border, attached to the other label) / LECTOTYPE Leptogaster datis Walker, 1849 by T. Dikow 2001 (red label)'. The specimen is double mounted (minuten in rectangular piece of plastic), is in poor condition (antennae, left meso leg, and meta legs broken; abdomen broken from the third segment on), and is deposited in the BMNH. The [male] holotype of Lasiocnemus pilipes is labelled 'Kwamouth, Congo 16[degrees]10'E 3[degrees]20'S. V.24.1915 [24.v.1915] / Lang & Chapin Collectors / Lasiocnemus Type pilipes [male] Curran No. (red label, handwritten except for 'Type' and 'No.')'. The specimen is directly mounted, is in good condition, and is deposited in the AMNH. Curran apparently mentioned additional type specimens (1 [female] allotype and 1 [female] paratype from Faradje as well as 1 [female] paratype from Garamba) in his description. These specimens could not be found in the AMNH collection and have not been examined. Type locality and distribution (Fig. 26): Sierra Leone, no exact locality. Cameroon, Central African Republic, DR Congo, Kenya, Nigeria, Senegal, Sierra Leone. Material examined: CAMEROON: 1 [male] Bossoum, 9[degrees]56'N 13[degrees]41'E, 16.v.1914; CENTRAL AFRICAN REPUBLIC: 1 [male] Fort Crampel, 6[degrees]59'N 19[degrees]11'E; DR CONGO: 1 [male] Eala, 0[degrees]04'N 18[degrees]17'E, xii.1934; 1 [male] Boma, 5[degrees]51'S 13[degrees]03'E; 1 [male] Albertville, 5[degrees]56'S 29[degrees]12'E, xii.1918; KENYA: 1 [male] Gelegele River, 0[degrees]53'S 35[degrees]02'E, vi.1913; NIGERIA: 1 [male] Samaru, 11[degrees]10'N 7[degrees]37'E, 21-29.vii.1970; 1 [male] Kateri, 9[degrees]42'N 7[degrees]27'E, 2.xii.1912 [genitalia slide examined]; SENEGAL: 1 [male] Mpak, 12[degrees]27'N 16[degrees]14'W, 8.xi.1977; SIERRA LEONE: 1 [male] Macdonald, 8[degrees]16'N 13[degrees]04'W, 25.xi.1993. Depository: BMNH, ISNB, MRAC, MZLU, ZMHB, ZSMC. Remarks: E. datis is the oldest described species of the datis species-group. The abdomen of the lectotype is apparently broken and therefore the [male] terminalia remained undescribed. Just before the end of this project I was able to investigate a single [male] specimen from Sierra Leone in which the type locality of datis is situated. The [male] terminalia of this specimen were indistinguishable from that of pilipes (of which some specimens had already been examined). Therefore, I am confident to delimit the boundaries of datis and consider the later described pilipes to be a junior synonym. Another [male] specimen from Senegal, also examined at the end of the project, supports the new synonymy by extending the distribution further west where no other species of the species-group has so far been found. Theodor (1976: 20) illustrated the [male] terminalia of this species, but labelled them artaphernes. [female] [female] specimens examined of representatives of the datis species-group: BENIN: 1 [female] Cotonou, 6[degrees]21'N 2[degrees]26'E, 29.v.1914; BURKINA FASO: 1 [female] Soumousso, 11[degrees]01'N 4[degrees]03'W, 11.x.1979; BURUNDI: 4 [female] [female] 2?? Rumonge, 3[degrees]58'S 29[degrees]26'E, 17.iv.1949, 21+22.i.1950; 1 [female] Minago, 3[degrees]47'S 29[degrees]21'E, 5.iv.1949; 1 [female] Mutonga, 3[degrees]23'S 30[degrees]29'E, 1.vi.1952; CAMEROON: 17 [female] [female] l? Bossoum, 9[degrees]56'N 13[degrees]41'E, 15, 16, 22, 23, 29.v., 1-10, 11-20.vi.1914; 1 [female] Limbe, 4[degrees]01'N9[degrees]11'S, 18.iii.1984; 1 [female] Kribi, 2[degrees]55'N9[degrees]54'E, 28-29.xi.1987; 1 [female] Campo, 2[degrees]22'N 9[degrees]49'E, 28-29.iii.1974; 1 [female] Johann-Albrechtshohe, iii.1896; CENTRAL AFRICAN REPUBLIC: 1 [female] Fort Crampel, 6[degrees]59'N 19[degrees]11'E; 1 [female] Dzanga-Sangha Reserve, 2[degrees]55'N 16[degrees]15'E, 6.vii.1998; DR CONGO: 1 [female] Eala, 0[degrees]04'N 18[degrees]17'E, 1.xi.1939; 1 [female] Bukavo, 2[degrees]30'S 28[degrees]51'E, 18.xii.1952; 1 [female] Kivu area, 3[degrees]03'S 28[degrees]06'E; 3 [female] [female] Boma, 5[degrees]51'S 13[degrees]03'E; 2 [female] [female] Lulua, 5[degrees]55'S 25[degrees]46'E, xii.1932,.iv.1933; 1 [female] Kasal; GHANA: 1 [female] Akosombo, 6[degrees]18'N 0[degrees]03'E, 22-25.vi.1973; 1 [female] Achimota, 5[degrees]37'N 0[degrees] 14'W, 12.x.1949; IVORY COAST: 1 [female] Yapi, 6[degrees]26'N 3[degrees]51'E, ix.1915; KENYA: 1 [female] Mt. Elgon, 1[degrees]06'N 34[degrees]34'E, 8-13.vi.1911; 2 [female] [female] Kuja Valley, 0[degrees]48'S 34[degrees]34'E, 30.iv.-1.v.1911; 1 [female] Masa Reserve, 1[degrees]26'S 34[degrees]55'E, 17.v.1913; 1? Athi, 2[degrees]15'S 38[degrees]03'E, vi.1941; 1 [female] Malindi, 3[degrees]13'S 40[degrees]07'E, v.1973; MALAWI: 2 [female] [female] 1? Senga Bay, 13[degrees]42'S 34[degrees]37'E, 7-8.iii.1987; NIGERIA: 2 [female] [female] Samaru, 11[degrees]10'N 7[degrees]37'E, 21-29.vii.1970; 1 [female] Baro, 8[degrees]36'N 6[degrees]25'E, 14-16.x.1910; 1? Onitsha, 6[degrees]09'N 6[degrees]46'E, vii.1910; 1? Lagos, 6[degrees]29'N 3[degrees]22'E, 3.xii.1956; 1 [female] Apapa, 7[degrees]12'N 4[degrees]01'E, 24.iii.1963; 1 [female] Mokwa, 9[degrees]16'N 5[degrees]02'E, 11-19.viii.1970; SENEGAL: 1 [female] Ziguinchor, 12[degrees]34'N 16[degrees]17'W, 13.viii.1979; SIERRA LEONE: 1 [female] 2?? Freetown, 8[degrees]29'N 13[degrees]14'W, 1940; 1? no additional data; TANZANIA: 1 [female] Ruo Valley, 10[degrees] 13'S 39[degrees]34'E, 5.iii.1913; 1 [female] NW Tanganjika; TOGO: 1 [female] Hinterland; UGANDA: 1 [female] Pakelle, 3[degrees]22'N 31[degrees]5 1'E, 17.xi.1958; 2 [female] Ngora, 2[degrees]06'N 33[degrees]26'E, 15-16.xi.1948; 1 [female] Serere, 1[degrees]31'N 33[degrees]26'E, 15.ix.1957; 1 [female] Buwalasi, 1[degrees]11'N 34[degrees] 4'E, 20-21.xi.1948; 1 [female] Nagongera, 0[degrees]46'N 34[degrees]00'E, 15.v.958; 1 [female] Kitega, 0[degrees]42'N 30[degrees]51'E, 11.vi.1950; 1 [female] Tororo Hills, 0[degrees]42'N 34[degrees]10'E, v.1951; 1 [female] Entebbe, 0[degrees]03'N 32[degrees]27'E, 28.x.1912; 1 [female] Masaka, 0[degrees]30'S 31[degrees]45'E, 1.xii.1948; 1? Bugalla Island, 0[degrees]38'S 32[degrees]20'E, v.1912; 1 [female] Madi county, 2[degrees]48'S 31[degrees]15'E, v.1927; COUNTRY UNKNOWN: 4 [female] [female] 1? no locality data. Depository: AMNH, BMNH, CUIC, ISNB, MRAC, NMSA, USNM, ZMHB, ZMUC, ZSMC. Distribution of [female] [female] of datis species-group (Fig. 26): Benin, Burkina Faso, Burundi, Cameroon, Central African Republic, DR Congo, Ghana, Ivory Coast, Kenya, Malawi, Nigeria, Senegal, Sierra Leone, Tanzania, Uganda. Euscelidia lata sp. n. (E. datis species-group) Figs 8C, 91, 26, 28A-C, K-L Etymology: Latin adjective latus = wide, broad; refers to the widened distal tip of the dorsal aedeagal sheath (Fig. 28L). Description: Head: Black; fc yellow pruinose, fc gib indistinct, mystax white, many macrosetae; prob and plp black, setae white; oc tr apruinose; occ silver pruinose, many white setae; Antennae--scp dark brown, white setae ventrally, white pruinose; ped brown, either white or black setae ventrally and black setae dorsally, white pruinose; pped brown, ventrally light brown, white pruinose; apsel hyaline. Thorax: Black; ppro peg large, silver pruinose; sct bluish-black, red spots antero-laterally, apruinose, margins silver pruinose, surface covered with long yellow setae, white setae on pruinose part; macrosetae: white, 1 npl s, 1 spal s; sctl silver pruinose, ds sctl s and sctl s long, white; Legs--orange and dark brown; pro fern orange, meso fem orange, antero-medially brown, meta fem clubbed, pale yellow proximally, proximal half of club black, distal half orange, club with erect yellow setae; pro tib light brown, pale yellow stripe anteriorly, meso tib brown, pale yellow stripe anteriorly, meta tib orange to black, pale yellow stripe not reaching tip anteriorly; first tar pale yellow, brown distad, remaining tar black, setae black; emp distinct, very short; Wings--densely covered with microtrichia, brown coloured; ptero distinct, brown; cell d terminating in 2 veins; hlt light brown. Abdomen: Orange to black; T predominantly black, anterior and posterior margins orange, yellow pruinose, T1 with many long yellow setae, T2 in proximal half with long yellow setae laterally, remaining T with short yellow and brown setae, S yellow pruinose, setae yellow; [male] terminalia Figs 28A-C, K-L--sur bent inwards distally; hypd distally broadened; d aed shea long, tip Fig. 28L, lat apod bifurcated. Type material--The [male] holotype is labelled 'COTE D'IVOIRE: Near Palmindustrie about 15 km West of Fresco 05[degrees]06'N 05[degrees]43'W JGH Londt 24.iv.1989 Airfield/Forest edge / HOLOTYPE Euscelidia lata sp. nov. det. T. Dikow 2001 (red label)'. The specimen is directly mounted, is in excellent condition, and is deposited in the NMSA. The 4 [female] [female] 4 [male] [male] paratypes have same labels as holotype except 'PARATYPE Euscelidia lata sp. nov. det. T. Dikow 2001 (yellow label)'. The specimens are directly mounted, are in good to excellent condition, and are deposited in the NMSA. Type locality and distribution (Fig. 26): Ivory Coast, Fresco (15 km W), 5[degrees]06'N 5[degrees]43'W. Cameroon, Ivory Coast, Liberia. Material examined: CAMEROON: 1 [female] Limbe, 4[degrees]01'N 9[degrees] 11'E, 17.iii.1984; 1 [male] Yaounde, 3[degrees]52'N 11[degrees]30'E, 5.xi.1987; 1 [male] Yaounde; 1 [male] Edea, 3[degrees]48'N 10[degrees]07'E, 31.xii.1977; 2 [male] [male] Kribi, 2[degrees]56'N 9[degrees]54'E, 19.iii.-7.iv.1974; 1 [male] Campo, 2[degrees]22'N 9[degrees]49'E, 28-29.iii.1974; IVORY COAST: 2 [female] [female] 3 [male] [male] Abidjan, 5[degrees]19'N 4[degrees]00'W, 21.iv.1989; LIBERIA: 1 [male] Bakratown (? = Baka, 6[degrees]11'N 8[degrees]47'W--plotted as such in Fig. 26), x.1926. Depository: MZLU, NMSA, USNM, ZMHB, ZSMC. Euseelidia milva sp. n. (E. datis species-group) Figs 25, 26, 28D-F, M Etymology: Latin noun milvus = kite, hawk; refers to the predatory habit of all Asilidae. Description: Head: Black; fc yellow pruinose, fc gib indistinct, mystax white, many macrosetae; prob and plp black, setae white; oc tr apruinose; occ silver pruinose, many white and yellow setae; Antennae--scp dark brown, white setae ventrally, white pruinose; ped brown, brown setae ventrally and dorsally, white pruinose; pped brown or orange, white pruinose; apsel hyaline. Thorax: Black; ppro peg large, silver pruinose; sct bluish-black, red spots antero-laterally, predominantly apruinose, margins silver pruinose, surface covered with long yellow setae, white setae on pruinose part; macrosetae: white, 1 npl s, 1 spal s; sctl silver pruinose, ds sctl s and sctl s long, white; Legs--orange to dark brown; pro fem orange, meso fern orange, antero-medially brown, meta fem clubbed, pale yellow proximally, proximal half of club black, distal half orange, club with erect yellow setae; pro tib light brown, pale yellow stripe anteriorly, meso tib brown, pale yellow stripe anteriorly, meta tib orange to black, pale yellow stripe not reaching tip anteriorly; first tar pale yellow, brown distad, remaining tar black, setae black; emp distinct, very short; Wings--densely covered with microtrichia, brown coloured; ptero distinct, brown; cell d terminating in 2 veins; hlt light brown. Abdomen: Orange to black; T predominantly black, anterior and posterior margins orange, yellow pruinose, T1 with many long yellow setae, T2 in proximal half with long yellow setae laterally, remaining T with short yellow and brown setae, S yellow pruinose, setae yellow; [male] terminalia Figs 28D-F, M--sur bent inwards distally; hypd enlarged proximally; d aed shea long, tip Fig. 28M, lat apod bifurcated. Type material--The [male] holotype is labelled 'W-Africa: Ghana; ca. 2 km NE' Hohoe: Bodenabbaugebiet [a place where soil is produced] BARKEMEYER (handwritten except for 'W-Africa: Ghana' and 'BARKEMEYER') 9.10.1998 [9.x.1998] (handwritten on backside of label) / HOLOTYPE Euscelidia milva sp. nov. det. T. Dikow 2001 (red label)'. The specimen is directly mounted, is in excellent condition ([male] terminalia attached to specimens pin in micro vial), and is deposited in the NMSA. A [male] paratype with the same locality data as the holotype, collected on 8.x.1998, and 'PARATYPE Euscelidia milva sp. nov. det. T. Dikow 2001 (yellow label)' is directly mounted, is in excellent condition, and is deposited in the NMSA. A [male] paratype is labelled 'W-Africa: Ghana; ca. 2 km NE' Hohoe: 'Farmland' am Folibli BARKEMEYER (handwritten except for 'W-Africa: Ghana' and 'Barkemeyer') 8.10.1998 [8.x.1998] (handwritten on backside of label) / PARATYPE Euscelidia milva sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is directly mounted, is in very good condition stylus of antennae broken), and is deposited in the NMSA. A [male] paratype is labelled 'a in cop. with b. (handwritten) / NIGERIA: N.W. State Mokwa. Zugurma. R. Eku. 11-19.viii.1970. P.H. Ward. B.M. 1970-604 / PARATYPE Euscelidia milva sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is double mounted (minuten in block of foam), is in very good condition (right meta leg broken), and is deposited in the BMNH. A [female] paratype is labelled 'b in cop. with a. (handwritten) / NIGERIA: N.W. State Mokwa. Zugurma. R. Eku. 11-19.viii.1970. P.H. Ward. B.M. 1970-604 / PARATYPE Euscelidia milva sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is double mounted (minuten in block of foam), is in good condition (met legs broken), and is deposited in the BMNH. A [male] paratype is labelled 'NIGERIA: N.W. State Mokwa. Zugurma. R. Eku. 11-19.viii.1970. P.H. Ward. B.M. 1970-604 / PARATYPE Euscelidia milva sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is double mounted (minuten in block of foam), is in good condition (right meso tib and meta leg broken), and is deposited in the BMNH. A [male] paratype is labelled 'UGANDA EAST MADI PARKELLE 17.11.58 [17.xi.1958] J. BOWDEN. / Euscelidia artaphernes Speiser det. H. Oldroyd, 1960. (handwritten except for 'det. H. Oldroyd, 196') / PARATYPE Euscelidia milva sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is double mounted (minuten in rectangular piece of plastic), is in very good condition ([male] terminalia attached to specimens pin in micro vial), and is deposited in the BMNH. Type locality and distribution (Fig. 26): Ghana, Hohoe, 7[degrees]09'N 0[degrees]27'E. Central African Republic, Ghana, Nigeria, Uganda. Material examined: CENTRAL AFRICAN REPUBLIC: 1 [male] Fort Crampel, 6[degrees]59'N 19[degrees]11'E. Depository: ZSMC. Euscelidia rapax Westwood, 1850 (E. datis species-group) Figs 27D-F, H Euscelidia rapax Westwood, 1850: 232; Hermann 1926: 147; Engel & Cuthbertson 1934: 36; Cuthbertson 1939: 136; Oldroyd 1939: 28; Janssens 1953: 5; Lindner 1955: 28; Hull 1962: 305; Oldroyd 1980: 357. Redescription: Head: Black; fc yellow pruinose, fc gib distinct, mystax yellow, many macrosetae, nearly as long as prob; oc tr apruinose; occ silver pruinose, setae yellow; Antennae (ped and pped broken)--scp brown, yellow setae ventrally; Thorax: Black; sct black, 2 orange spots antero-laterally, predominantly apruinose, margins white pruinose, yellow setae on anterior, lateral, and posterior margins, leaving dorsal surface nearly uncovered; sctl white pruinose, ds sctl s and sctl s long, yellow; Legs--red-orange to brown; pro and meso fern light brown, meso fem with brown ring medially, meta fern pale yellow proximally, brown medially, orange distad, clubbed, club densely covered with long, erect yellow setae; pro and meso tib brown, yellow stripe anteriorly, meta tib orange to light brown, yellow stripe not reaching tip anteriorly; emp distinct, very short; Wings--densely covered with microtrichia, brown coloured; ptero indistinct, brown; cell d terminating in 2 veins; hlt light brown; Abdomen: Brown; T on anterior and posterior margins orange, grey pruinose, setae yellow, T1 with long yellow setae, T2 in proximal half with yellow setae laterally; [male] terminalia Figs 27D-F, H--sur bent inwards distally; hypd proximally greatly enlarged, with rectangular membranous area ventrally; d aed shea long, tip broken, lat apod bifurcated. Type material--The [male] lectotype, here designated to preserve stability and make more universal the use of this name, is labelled 'Euscelidia rapax Westw Trans Ent Soc Vol 5 pl 23 Afr. trop?. (handwritten) / W / WESTWOOD'S TYPE OF Euscelidia rapax Tr Ent soc Vol V p232 MS. (1902) by J. W. Yerbury (species name and literature citation handwritten) / Asil. 1. (handwritten) /TYPE Dip.: 152 Euscelidia rapax Westwood Hope Dept. Oxford (species name and type no. handwritten) / LECTOTYPE Euscelidia rapax Westwood, 1850 by T. Dikow 2001 (red label)'. The specimen is double mounted (minuten in rectangular piece of cardboard), is in good condition (left pro leg broken, abdomen broken, but attached to piece of cardboard), and is deposited in the OXUM. Type locality and distribution: 'Africa tropicali?', no exact locality, distribution uncertain. Remarks: No additional specimens of this species have been examined. The tip of the dorsal aedeagal sheath is apparently broken in the lectotype. The [male] terminalia of all other species in the species-group, however, cannot be related to the terminalia found in rapax. Euscelidia discors Speiser, 1913 Euscelidiadiscors Speiser, 1913: 141; Hermann 1926: 147; Hull 1962: 305; Oldroyd 1980: 356. Type material--The 1 [female] 3 [male] [male] syntypes have to be considered lost. I was unable to locate the depository for the type specimens. The concept of discors remains unclear and no neotype designation is possible at this stage. Type locality (Fig. 16): Cameroon, Douala, 4[degrees]02'N 9[degrees]42'E. Remarks: Speiser's description is very poor in providing information that is useful for the identification of the species. Generally, many species would fit that description. There is a specimen deposited in the ZSMC that was identified as E. discors by E. O. Engel, but it is a representative of E. procula--this might direct to a probable synonymy of the two species withprocula then being the senior synonym. Although I examined many specimens of Euscelidia from Cameroon I am not in the position to make further comments about this species. The type locality is plotted in Fig. 16. [FIGURE 29 OMITTED] Euscelidia dorata Oldroyd, 1970 Figs 29, 30 Euscelidia dorata Oldroyd, 1970:220 1980: 356. Diagnosis: The species is distinguished from congeners by the entirely pruinose sct, only four setae comprising the mystax, vein [R.sub.4] bulging anteriorly constricting cell [r.sub.2+3], and features of the [male] terminalia (very long d aed shea). Redescription: Head: Black; fc silver pruinose, fc gib indistinct, mystax white, 4 macrosetae, as long or longer than prob; oc tr yellow pminose dorsally; occ yellow pruinose, few white setae; Antennae--scp and ped brown, setae dark brown; pped brown, white pminose; apsel brown; Thorax: Brown; sct hump-backed, red-brown, entirely covered with white pminosity, 3 indistinct median stripes shine through the pruinosity, macrosetae: yellow, 1 npl s, 1 spal s; very few setae on anepst directing anteriorly; sctl silver pruinose, ds sctl s and sctl s very short; Legs--yellow to brown; fern light brown, meta fern yellow proximally, clubbed; pro and meso tib yellowish-brown, meta tib brown; emp long, longer than half of length of clw; Wings--microtrichia densely arranged on remigium; ptero distinct, brown; vein [R.sub.4] bulging anteriorly constricting cell [r.sub.2+3], cell d terminating in 3 veins; hlt yellow; Abdomen: yellow or brown; T white pminose, T1 with white setae, remaining T with short white and brown setae, S white pminose; [male] terminalia Fig. 30--sur pointed distally, lobe ventrally; hypd ventral margin straight; gonst large, prominent structure arising from proximal hypd; d aed shea tubular, very long (extending [male] terminalia); lat apod simple. [FIGURE 30 OMITTED] Type material--The [male] holotype is labelled 'HOLOTYPUS (red label with black submarginal border) / Congo Belge, P. N. G. [Parc National Garamba] Miss. H. De Saeger II/fd/15, 22-IX-1951 Rec. H. De Saeger. 2464 / Euscelidia dorata Oldroyd det. H. Oldroyd 1966 HOLOTYPE (handwritten except for 'det. H. Oldroyd 19')'. The specimen is double mounted (minuten in block of foam), is in very good condition (right pro leg broken; [male] terminalia attached to specimens pin in micro vial), and is deposited in the MRAC. The [female] paratype is labelled 'Congo Belge, P.N.G. Miss H. De Saeger II/gd/11, 12-VII-1951 Rec. H. De Saeger. 2071 / Euscelidia dorata Oldr. det, H. Oldroyd 1966. (handwritten except for 'det. H. Oldroyd 1966') / Paratype (red label)'. The specimen is double mounted (minuten in block of foam), is in fairly poor condition (antennae, right pro leg, left meso and meta legs, and abdomen broken), and is deposited in the ISNB. The [male] paratype is labelled 'Congo Belge, P.N.G. Miss H. De Saeger II/fe/Gar, 16-II-1951 Rec. J. Verschuren. 1269 / Paratype (red label) / Euscelidia dorata Oldr. det. H. Oldroyd 1966. (handwritten except for 'det. H. Oldroyd 1966')'. The specimen is double mounted (minuten in block of foam), is in fairly good condition (antennae, left pro leg, and all right legs broken), and is deposited in the ISNB. Type locality and distribution (Fig. 29): DR Congo, Garamba National Park, 4[degrees]06'N 29[degrees]27'E. Angola, DR Congo, Ethiopia, Tanzania, The Gambia, Zambia, Zimbabwe. Material examined: ANGOLA: 1 [female] Nova Lisboa, 12[degrees]45'S 15[degrees]43'E, 5-7.iii.1972; 1 [female] Alto Hama, 12[degrees] 14'S 15[degrees]33'E, 7.iii.1972; DR CONGO: 11 [female] [female] 1 [male] 1? Garamba National Park, 4[degrees]06'N 29[degrees]27'E, 15.ii.1950, 17.ii., 27.iv., 30.vi., 22.ix., 23.xi., 10.xii., 20.xii., 26.xii.1951; ETHIOPIA: 1 [female] Bahir Dar, 11[degrees]33'N 37[degrees]21'E, 11.x.1968; 1 [female] Adowa, 14[degrees]10'N 38[degrees]54'E, 18.x.1957; TANZANIA: 1 [female] Kafukola, 8[degrees]04'S 31[degrees]57'E, 12.i.1956; THE GAMBIA: 1 [female] Bakau, 13[degrees]29'N 16[degrees]41'W, 23.xi.1977; ZAMBIA: 1 [male] Kasanka National Game Reserve, 12[degrees]30'S 30[degrees] 15'E, 9-21 .xii.1989; ZIMBABWE: 1? Harare, 17[degrees]49'S 31 [degrees]02'E, 18.vi.1938. Depository: BMNH, ISNB, MRAC, MZLU, NMSA, SMNS, USNM. Euscelidia erichthenii sp. n. Figs 31, 32, 61 Etymology: The specific name 'erichthenii' is a noun in apposition. The name is proposed by the Gesellschafl fur Biologische Systematik (Organisms, Diversity & Evolution) to acknowledge the contribution to the understanding of the evolution of Vertebrata made over many years by the palaeontologist Prof. Dr Erich Thenius, Vienna, Austria, who has been awarded an honorary membership of the society. Diagnosis: The species is distinguished from congeners by the predominantly apruinose sct and features of the d terminalia (hypd bent upwards, divided into 2 lobes, d aed shea short, blunt). Description: Head: Black; fc yellow pruinose, sometimes silver pruinose on lower facial margin, fc gib indistinct, mystax white, many macrosetae; prob and plp brownish-black with white setae; oc tr silver pruinose dorsally; occ silver pruinose, setae white; Antennae--scp dark brown, white setae ventrally; ped dark brown, brown setae ventrally and dorsally, white pruinose; pped dark brown, white pruinose; apsel dark brown. Thorax: Black; ppro peg small, distinct, silver pruinose; anterior margin of sct yellow pruinose, lateral and posterior margins silver pruinose, 3 narrow median yellow pruinose stripes on sct, 2 lateral stripes sometimes reaching posterior margin, short yellow or white setae on pruinose part; macrosetae: 1 white npl s, 1 black spal s; sctl white pruinose, ds sctl s and sctl s short, white; Legs--yellow to brown; fern yellow proximally, brown distad, meta fern slightly clubbed, pale yellow stripe not reaching club dorsally, narrow longitudinal dark brown stripe laterally, setae white; tib brown, pale yellow stripe not reaching tip on meta tib anteriorly; first tar yellow proximally, brown distad, remaining tar brown, setae black; emp minute; Wings--hyaline, but microtrichia scattered on remigium and distal margin; ptero distinct, brown; cell d terminating in 2 veins; hlt light brown. Abdomen: Black; T brown pruinose dorsally, grey pruinose laterally, T1 with long brown setae laterally, T2 in proximal half with white setae laterally, remaining T with yellow and brown setae, S grey pruinose; [male] terminalia Figs 31-32--sur pointed distally, lobe ventrally; hypd bent upwards, divided into 2 large lobes; d aed shea short, blunt; lat apod simple. Type material--The d holotype is labelled 'ETHIOPIA Bahar Dar 12.10.1968 K.W. u. H. Harde leg (blue label) / Euscelidia spec. det. Miksch (handwritten except for 'det. Miksch') / HOLOTYPE Euscelidia erichthenii sp. nov. det. T. Dikow 2001 (red label)'. The specimen is double mounted (minuten in rectangular piece of cardboard), is in very good condition (right meta leg broken; d terminalia attached to specimens pin in micro vial), and is deposited in the SMNS. The 2rid paratypes have same labels as holotype except 'PARATYPE Euscelidia erichthenii sp. nov. det. T. Dikow 2001 (yellow label)'. The specimens are double mounted (minuten in rectangular piece of cardboard), are in very good condition (ld with right meta leg and pped broken), and are deposited in the SMNS. [FIGURE 31 OMITTED] [FIGURE 32 OMITTED] Type locality and distribution (Fig. 61): Ethiopia, Bahar Dar (= Bahir Dar), 11[degrees]33'N 37[degrees]21'E. Ethiopia. Material examined: 1 [female] without locality label (SMNS) [probably caught at the same locality as type material]. Euscelidia fastigium Martin, 1964 Fig. 16 Euscelidiafastigium Martin, 1964: 276; Oldroyd 1980: 356. Diagnosis: The species is distinguished from congeners by the entirely pruinose sct, only four setae comprising the mystax, and its distribution on Madagascar. Redescription: Head: Black; fc silver pminose, fc gib indistinct, mystax white, 4 macrosetae; oc tr yellow pminose dorsally; occ grey pminose, few white setae; Antennae--scp and ped brown, setae dark brown; pped brown, white pminose; apsel brown; Thorax: Brown; sct hump-backed, entirely covered with white and light brown pruinosity, macrosetae: yellow, 1 npl s, 1 spal s; very few setae on anepst directing anteriorly; sctl silver pruinose, ds sctl s and sctl s very short; Legs--yellow to brown; fern light brown, meta fern yellow proximally, clubbed; tib light brown to brown, yellow stripe anteriorly, not reaching tip on meta fib; emp long, longer than half of length of clw; Wings--microtrichia evenly scattered on remigium; ptero distinct, brown; cell d terminating in 3 veins; hlt yellow; Abdomen: brown; T white pruinose, T1 with white setae, remaining T with short white and brown setae, S white pruinose; cy terminalia--crcr specimens unknown. Type material--The [male] holotype is labelled 'HOLOTYPE Euscelidiafastigium Chas. H. Martin (species name handwritten, red label) / MADAGASCAR Maj. [Majunga Province] Ambato-Boeni 23.VI.58 [23.vi.1958] F. Keiser (red label) /Euscelidia fastigium Martin (handwritten, yellow label with black submarginal border)'. The specimen is double mounted (minuten in triangular piece of cardboard), is in very good condition (right pro and left meso leg broken), and is deposited in the NHMB. Type locality and distribution (Fig. 16): Madagascar, Ambato Boeny, 16[degrees]28'S 46[degrees]42'E. Madagascar. Remarks: The species is apparently known only from the single type specimen. I was able to investigate only three additional specimens of Euscelidia from Madagascar that are clearly not belonging tofastigium. The poor condition of the material and the lack of male specimens decided me against describing these specimens as a new species. The label data are as follows: 1 [female] Manjakatompo, 19[degrees]19'S 47[degrees]25'E, l.xi.1948 (BMNH); 2?? Andringitra-Ambalavao, Plateau Soaindrana, 21[degrees]31'S 47[degrees]02'E, 14-17.i.1958 (NMSA). Euscelidia festiva Janssens, 1954 Figs 33A-E, 61 Euscelidia festiva Janssens, 1954a: 123 1954b: 400; Oldroyd 1960:409 1980: 356. Diagnosis: The species is distinguished from congeners by the predominantly apruinose sct, the densely arranged microtrichia on the wings, and the yellow legs. Redescriptions: Head: Black; fc silver pruinose, fc gib indistinct, mystax white, 10-12 macrosetae, sometimes nearly as long as prob; oc tr apruinose; occ yellow pruinose dorsally, silver pruinose laterally, setae white; Antennae--scp and ped brown, setae brown; pped brown, white pruinose; apsel dark brown; Thorax: Black; sct predominantly apruinose, anterior, lateral, and posterior margins yellow pruinose, yellow setae on pruinose part, dc setae sometimes distinct, brown; sctl silver pruinose, ds sctl s and sctl s long, yellow; Legs--yellow; meta fem pale yellow proximally, clubbed, setae white; meta tib yellow proximally, brown distad; emp minute; Wings--densely covered with microtrichia; ptero distinct, brown; cell d terminating in 3 veins; hlt light brown; Abdomen: Black; T2-3 and proximal half of T4 predominantly apruinose, grey pruinose laterally, remaining T grey pruinose, S2 apruinose, remaining S grey pruinose; terminalia Figs 33A--E--sur pointed distally, large lobe ventrally; hypd bent upwards, enlarged distally; d aed shea short, blunt, lat pr aed short; lat apod simple. [FIGURE 33 OMITTED] Type material--The holotype of undeterminable gender is labelled 'HOLOTYPUS (red label with black submarginal border) / TYPE (red label with black submarginal border) / Congo beige: E N. U. [Parc National de l'Upemba] Kafwi af. dr. Lufwa [riviere Kafwi, affluente droite Lufwa] (1.780 m.) 5-III-1948 Mis. G. E de Witte. 1375a / E. Janssens det., 1952 Euscelidia festiva Em. Janssens (handwritten except for 'E. Janssens det., 19')'. The specimen is double mounted (minuten in block of foam), is in fairly good condition (antennae, meta legs, and terminal segments of abdomen broken), and is deposited in the MRAC. 21 paratypes (11 [female] [female] 6[male][male]4?) from the Upemba National Park are deposited in the ISNB. The following list will only provide the locality data, date of collection, and collector: 5[female] [female] 3[male][male] 2?? Mukana, 1810 m, 4+6+18.iii.1948, G.F. de Witte; 1[female] 1? Grande Kafwe, 1780 m, 5.iii.1948, G.F. de Witte; 3 [female] [female] 1[male] 1? River Kafwiaf, 1780 m, 5.iii. 1948, G.F. de Witte; 1 [female] 2[male][male] Kabwekanono, 1815m, 6+8.iii. 1948, G.F. de Witte; 1 [female] Lusinga, 13.vi.1945, G.F. de Witte. Two additional paratypes apparently belong not to E. festiva, but are species of Leptogaster. Type locality and distribution (Fig. 61): DR Congo, Upemba National Park, Lufwa, 9[degrees]40'S 27[degrees] 12'E. Cameroon, DR Congo, Malawi, Nigeria, Uganda, Zambia. Material examined: CAMEROON: 2[female] [female] Bossoum, 9[degrees]56'N 13[degrees]41'E, 1-10.v.1914; 1 [female] 1[male] Kanjang, 6.v. 1909; DR CONGO: 1 [female] 2[male][male] Garamba National Park, 4[degrees]06'N 29[degrees]27'E, 18.v.1950, 27.ix.1950, 11.vi.1952; 1? Kamina [several such localities], 1.iii.1960; MALAWI: 1 [female] Mulanje, 15[degrees]57'S 35[degrees]37'E, 13.i.1913; NIGERIA: 1[male] Samaru, 11[degrees]10'N 7[degrees]37'E, 24.vi.1968; UGANDA: l? Abi Farm, 3[degrees]05'N 30[degrees]56'E, 12.xi.1958; 1[male] 1? Mbale-Sipi, 1[degrees]05'N 34[degrees]09'E, 14.v.1957; ZAMBIA: 1[male] Kasanka National Game Reserve, 12[degrees]30'S 30[degrees]15'E, 16-22.xii.1989; COUNTRY UNKNOWN: 1? no locality data. Depository: BMNH, ISNB, MRAC, NMSA, ZMHB. Remarks: Additionally to the specimens examined, Oldroyd (1960: 409) listed the species from Handeni (5[degrees]26'S 38[degrees]01'E), Tanzania. Euscelidia fistula sp. n. Figs 14, 33F-K Etymology: Latin noun fistula = tube, pipe; refers to the tubular dorsal aedeagal sheath. Diagnosis: The species is distinguished from congeners by the predominantly apruinose sct, the densely arranged microtrichia on the wings, and features of the [male] terminalia (d aed shea tubular, hypd bent upwards). Description: Head: Black; fc silver pruinose, fc gib indistinct, mystax white, many macrosetae; prob black, setae white, plp brown, setae brown; oc tr apruinose; occ silver pruinose, setae white and yellowish; Antennae--scp brown, white setae ventrally, white pruinose; ped brown, brown setae ventrally and dorsally, white pruinose; pped brown, white pruinose; apsel brown. Thorax: Black; ppro peg large, silver pruinose; sct predominantly apruinose, silver pruinosity restricted to anterior, lateral, and posterior margins (sometimes antero-lateral margin apruinose), white setae on pruinose part, yellow setae scattered on apruinose surface; presut dc setae sometimes distinct, macrosetae: 1 white or black npl s, 1 black spal s; sctl white pruinose, ds sctl s and sctl s long, yellow or brown; Legs--yellow to brown; fern light brown, meta fem slightly clubbed, pale yellow proximally, narrow longitudinal dark brown to black stripe laterally, setae white; pro and meso tib light brown, pale yellow stripe anteriorly, meta tib light brown, pale yellow stripe not reaching tip anteriorly, black longitudinal stripes laterally; tar 1-3 light brown, brown distad, remaining tar brown, setae black; emp minute; Wings--densely covered with microtrichia; ptero distinct, brown; cell d terminating in 2 veins; hlt light brown. Abdomen: Black; T sometimes orange on anterior and posterior margins, predominantly brown pruinose, anterior, lateral, and posterior margins narrowly grey pruinose, T1 with long yellow and brown setae laterally, T2 with white setae laterally, remaining T with yellow and brown setae, S grey pruinose; [male] terminalia Figs 33F-K--sur pointed distally, lobe ventrally; hypd bent upwards, divided into 2 short lobes; d aed shea tubular, medium length; lat apod simple. Type material--The [male] holotype is labelled 'Johannesburg Transvaal leg. Zumpt / E. Janssens det., 1956 Euscelidia festiva E. Janssens (handwritten except for 'E. Janssens det., 195') / HOLOTYPE Euscelidia fistula sp. nov. det. T. Dikow 2001 (red label)'. The specimen is double mounted (attached to rectangular piece of cardboard), is in very good condition ([male] terminalia attached to specimens pin in micro vial), and is deposited in the ISNB. A [female] paratype is labelled 'SOUTH AFRICA 2527CC Transvaal nr. Derby 3.II.1978 JGH. Londt Open grassveld area / PARATYPE Euscelidia fistula sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is directly mounted, is in excellent condition, and is deposited in the NMSA. A (2 paratype is labelled 'SUID-AFRIKA Potch [Potchefstroom] 7-1-1943 H.P.V. HEERDEN ('Potch' and date handwritten) / PARATYPE Euscelidia fistula sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is double mounted (minuten in rectangular piece of cardboard), is in very good condition, and is deposited in the NMSA. A paratype of undeterminable gender is labelled 'PRETORIA 13.II. 1957 L. Vari (date handwritten) / collection TRANSVAAL MUSEUM (blue label) / Euscelidia sp. det. H. Oldroyd 1973 (handwritten except for 'det. H. Oldroyd 19') / PARATYPE Euscelidia fistula sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is directly mounted, is in good condition (left meta leg and abdomen from third segment broken), and is deposited in the NMSA. A [female] paratype is labelled 'ZIMBABWE: 2028AD Matopos National Park Jan. 1977 Coll: Mr Fox / PARATYPE Euscelidia fistula sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is directly mounted, is in relatively poor condition (left meta leg broken, abdomen damaged, specimen greasy), and is deposited in the NMSA. Type locality and distribution (Fig. 14): South Africa, Gauteng Province, Johannesburg, 26[degrees] 12'S 28[degrees]02'E. South Africa, Zimbabwe. Material examined: SOUTH AFRICA: Northern Prov: 1? Marico, 24[degrees]13'S 26[degrees]54'E, ii. 1918 [specimen in very poor condition]. Depository: BMNH. Euscelidia flava sp. n. Figs 34, 36 Etymology: Latin adjective flavus = golden-yellow; refers to the yellow pruinose scutum. Diagnosis: The species is distinguished from congeners by the predominantly yellow pruinose set and the densely arranged microtrichia on the wings. Description: Head: Black; fc silver or light yellowish pruinose, fc gib indistinct, mystax white, many macrosetae; prob and plp brown, setae white; oc tr apruinose dorsally; occ silver pruinose, setae white; Antennae--scp light brown, brown setae dorsally, white pruinose; ped light brown, brown setae ventrally and dorsally, white pruinose; pped light brown, white pruinose; apsel brown. Thorax: Brown to black; ppro peg small, distinct, silver pruinose; sct antero-laterally brown, predominantly yellow pruinose, indistinct apruinose spots laterally; macrosetae: 1 white npl s, 1 black spal s; sctl yellow pruinose, ds sctl s and sctl s long, yellow; Legs--yellow to brown; fern light brown to brown, meta fern pale yellow proximally, clubbed; pro tib yellow, meso and meta tib brown, pale yellow stripe not reaching tip anteriorly; first tar yellow, brown distad, remaining tar brown, setae black; emp about a fourth of length of clw; Wings--densely covered with microtrichia; ptero distinct, brown; cell d terminating in 2 veins; hlt light brown. Abdomen: Brown; yellow pruinose, T1 with long white setae laterally, T2 in proximal half with white setae laterally, remaining T with short white and yellow setae, S yellow pruinose; [male] terminalia Fig. 34--sur not sharply pointed distally, lobe ventrally; hypd with pointed lobes distally; d aed shea medium length; lat apod bifurcated. Type material--The [male] holotype is labelled 'Ceylon [Sri Lanka], N. Prov. [Northern Province] 2 mls [3,2 km] E Paraiyanalankulam 20 mls [32 km] W Vavuniya 15.II.62. [15.ii.1962] Loc. 62 / Swept in dry forest / Lund University Ceylon Expedition 1962 Brinck-Andersson-Cederholm / HOLOTYPE Euscelidia flava sp. nov. det. T. Dikow 2001 (red label)'. The specimen is directly mounted, is in excellent condition, and is deposited in the MZLU. A [male] paratype with the same label information, except for 'PARATYPE Euscelidia flava sp. nov. det. T. Dikow 2001 (yellow label)' is directly mounted, in relatively poor condition (antennae, prob, right pro tib, left meso leg, and right meso tarsus broken; [male] terminalia attached to specimens pin in micro vial), and is deposited in the MZLU. A [female] paratype is labelled 'Ceylon, NW. Prov [North Western Province] 7 mls [11,2 km] NE Puttalam 1.II.62. [1.ii. 1962] Loc. 43 / Swept on dry grassland /Lund University Ceylon Expedition 1962 Brinck-Andersson-Cederholm / PARATYPE Euscelidia flava sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is directly mounted, is in excellent condition, and is deposited in the MZLU. A [male] paratype and a paratype of undeterminable gender are labelled 'Ceylon, N Prov [Northern Province] Small stream 2 mls [3,2 km] E Mankulam 14.II.62. [14.ii.1962] Loc. 75 / Dry meadow [/ specimen] and Swept along roads in dry forest [undeterminable gender specimen] /Lund University Ceylon Expedition 1962 Brinck-Andersson-Cederholm / PARATYPE Euscelidia flava sp. nov. det. T. Dikow 2001 (yellow label)'. Both specimens are directly mounted, are in excellent condition (except for broken abdomen in one specimen), and are deposited in the MZLU. [FIGURE 34 OMITTED] Type locality and distribution (Fig. 36): Sri Lanka, Paraiyanalankulan, 8[degrees]46'N 80[degrees]10'E. Sri Lanka. Material examined: Sri Lanka: 1[male] Ma Villu, 8[degrees]42'N 80[degrees]02'E, 17-21.ii.1979. 1 [female] Kafukola, 8[degrees]04'S 31[degrees]57'E; 1 [female] Gaikadawala, 13.iii. 1976. Depository: USNM. Euscelidiafrancoisi (Janssens, 1953), comb. n. Figs 35, 46 Dolichoscius francoisi Janssens, 1953: 2: Hull 1962: 318; Oldroyd 1980: 356. Dolichoscius longipes Janssens, 1954a: 120; Oldroyd 1980: 356. syn. n. Diagnosis: The species is distinguished from congeners by the pale yellow wing veins C, Sc, and [R.sub.1], the narrow wing cell [a.sub.2], and the tuft of setae on the meta tib and tarsi. Redescription: Head: Black; fc yellow pruinose, fc gib indistinct, mystax white, many macrosetae; oc tr silver pruinose dorsally; occ silver pruinose, setae white; Antennae--scp brown, brown setae ventrally, white pruinose; ped dark brown, brown setae ventrally and dorsally, white pruinose; pped brown, white pruinose proximally and ventrally; apsel hyaline; Thorax: Black; ppro peg small, distinct, silver pruinose; sct predominantly apruinose, bluish-black, anterior, lateral, and posterior margins silver pruinose, long white setae on pruinose part, presut and psut dc setae long, macrosetae: white, 1 npl s, 1 spal s; sctl silver pruinose, ds sctl s and sctl s long, white; Legs--yellow to brown; pro fern light brown, meso fern light brown in proximal half, distal half brown, meta fern pale yellow proximally, clubbed, club brown; tib brown, yellow stripe not reaching tip anteriorly, distal half of meta tib and tar 1-4 with long brown setae laterally; emp minute; Wings--hyaline, very few microtrichia on remigium, veins yellow in proximal half of wing, brown distally, C, Sc, and [R.sub.1] completely pale yellow; ptero indistinct; cell d terminating in 3 veins; hlt yellow; Abdomen: Black; T predominantly brown pruinose, lateral margin grey pruinose, T1 with long white setae laterally, T2 in proximal third with white setae laterally, remaining T with short yellow and brown setae, S grey pruinose; [male] terminalia Fig. 35--sur pointed distally, lobe ventrally; hypd bent upwards, divided into 2 lobes; d aed shea tubular, medium length; lat apod simple. Type material--The [male] holotype of Dolichoscius francoisi is labelled 'Urundi: Terr. de Muhinga 11.V. 1952 EL. Francois ('Muhinga' and date handwritten) / Colline: Muhinga (Butezana) alt. 1700 m. ('Muhinga', 'Butezana', and '1700' handwritten)/E. Janssens det., 1953 Dolichoscius Francoisi n. sp. (species name and '53' handwritten) /TYPE (red label with black border around 'TYPE') / cf. Bull. Inst. Sc. Nat. Belg. XXIX, 1953, no 42 p. 2, figs. 1-2 (citation handwritten) / R. I. Sc. N. B. I. G. 20.102 (numbers handwritten) /Euscelidiafrancoisi (Janssens, 1953) det. T. Dikow 2001'. The specimen is double mounted (minuten in block of foam), is in very good condition (right meta leg broken), and is deposited in the ISNB. [FIGURE 35 OMITTED] Two [female] [female] and one [male] paratypes have same collection details as holotype. The specimens are double mounted (minuten in block of foam), are in very good condition, and are deposited in the ISNB. Two [female] [female] paratypes are labelled 'Urundi: Terr. de Bururi 9.II. 1952 F.J. Francois ('Bururi' and date handwritten) / Colline: Bururi (Burursi) alt. 1900 m. ('Bururi', 'Burursi', and '1900' handwritten) / Paratype (red label with black border around 'Paratype') / Euscelidia francoisi (Janssens, 1953) det. T. Dikow 2001'. The specimens are double mounted (minuten in block of foam), are in very good condition, and are deposited in the ISNB. The holotype of undeterminable gender of Dolichoscius longipes is labelled 'HOLOTYPUS (red label with black submarginal border) / TYPE (red label, black border around 'TYPE') / CONGO beige: P.N.U. Mukana (1.810 m.) 14-IV- 1947 Mis. G.F. de Witte. 275a / COLL. Mus. Tervuren (black border) / E. Janssens det., 1951 Dolichoscius longipes E. Janss. (handwritten except for 'E. Janssens det., 19') / Euscelidia francoisi (Janssens, 1953) det. T. Dikow 2001'. The specimen is double mounted (minuten in block of foam), is in poor condition (right pped, fight pro tib, right meso tarsus, left meta leg, and abdomen from fourth segment broken), and is deposited in the MRAC. Type locality and distribution (Fig. 46): Burundi, Muhinga, 2[degrees]50'S 30[degrees]20'E, Burundi, DR Congo, Mozambique, Rwanda. [FIGURE 36 OMITTED] Material examined: DR CONGO: 1 ? Mukana, 9[degrees] 15'S 27[degrees] 12'E, 14.iv. 1947; 1 [female] Kasende, vii.1934; MOZAMBIQUE: 1 [female] Maputo, 25[degrees]57'S 32[degrees]34'E, iv.--vii.1914; RWANDA: 2[male][male] Kibungu, 2[degrees]10'S 30[degrees]32'E, 2.ii.1953. Depository: BMNH, ISNB, MRAC. Euscelidia glabra sp. n. Figs 37, 47 Etymology: Latin adjective glaber = hairless; refers to the apruinose, bare scutum. Diagnosis: The species is distinguished from congeners by the predominantly apruinose sct, the few microtrichia on the wings, and features of the U terminalia (shape of aed). Description: Head: Black; fc yellow pruinose, fc gib distinct, mystax white, many macrosetae; prob and plp brownish-black, setae white; oc tr apruinose; occ silver pruinose, setae white; Antennae--scp brown, brown setae ventrally; ped brown, brown setae ventrally and dorsally, white pruinose; pped brown, white pruinose; apsel brown. Thorax: Black; ppro peg small, distinct, silver pruinose; sct bluish-black, predominantly apruinose, margins silver pruinose, white setae antero-laterally and on margins, presut and psut dc setae present; macrosetae: white, 1 npl s, 1 spal s; sctl silver pruinose, ds sctl s and sctl s long, white; Legs--yellow to brown; pro fern yellow, meso fern yellow proximally, remaining part brown, meta fern in proximal half pale yellow, clubbed, club brown, setae white; rib yellow proximally, brown in remaining part, pale yellow stripe not reaching tip on meta tib anteriorly; tar brown, black setae; emp short, about a fifth of length of clw; Wings--hyaline, but microtrichia evenly scattered on remigium and distal margin; ptero distinct, brown; cell d terminating in 3 veins; hlt light brown. Abdomen: Black; T predominantly brown pruinose, anterior margin grey pruinose laterally, posterior margin grey pruinose, T2 in proximal half with apruinose spot, T1 with long white setae laterally, T2 in proximal half with white setae laterally, remaining T with white and yellow setae, S grey pruinose; [male] terminalia Fig. 37--sur pointed distally, lobe ventrally; hypd bent upwards, pointed distally; d aed shea short, blunt, paired appendage developed ventrally; lat apod simple. Type material--The [male] holotype is labelled 'S. India: Karnataka. Gersoppa (Jog Falls) 19-24.xi. 1977 c. 600 m Zool. Mus. Copenhagen Exp. / HOLOTYPE Euscelidia glabra sp. nov. det. T. Dikow 2001 (red label)'. The specimen is directly mounted, is in excellent condition, and is deposited in the ZMUC. 17 [female] [female] 9[male][male] paratypes have same labels as holotype except 'PARATYPE Euscelidia glabra sp. nov. det. T. Dikow 2001 (yellow label)'. The specimens are either directly mounted or double mounted (minuten in block of foam), are in good to excellent condition, and are deposited in the ZMUC. Type locality and distribution (Fig. 47): India, Karnataka Province, Gersoppa, 14[degrees] 15'N 74[degrees]39'E. India. Euscelidia gutianensis Shi, 1995 Euscelidia gutianensis Shi, 1995: 228. Type material--The [female] holotype is deposited in the IZAS. The specimen has not been examined. Type locality: China, Zhejiang Province, Mt. Gutian, 29[degrees] 18'N 118[degrees] 12'E. [FIGURE 37 OMITTED] Remarks: Two features given in the short English description might be useful to distinguish the species from others. These are (1) 'scutum yellow with 3 black longitudinal stripes dorsally' and (2) 'hind femora with wide black ring near apex'. I have not examined material of this species and therefore no further comments about this species can be made. The type locality is plotted in Fig. 1. Euscelidia hesperia sp. n. Figs 29, 38 Etymology: Latin adjective hesperius = western; refers to the distribution in western Africa. Diagnosis: The species is distinguished from congeners by the predominantly apruinose sct, the distinct alveoli on the sct, and the densely arranged microtrichia on the wings. Description: Head: Black; fc silver pruinose, fc gib indistinct, mystax white, many macrosetae; prob and pip brown-black, setae white; oc tr white pruinose dorsally; occ silver pruinose, setae white; Antennae--scp dark brown, brown setae ventrally, white pruinose; ped dark brown, brown setae ventrally and dorsally, white pruinose; pped brown, white pruinose proximally and ventrally; apsel brown. Thorax: Black; ppro peg small, distinct, silver pruinose; sct predominantly apruinose, grey pruinosity restricted to anterior, lateral, and posterior margins, yellow setae with distinct alveoli scattered on surface, leaving 2 narrow, median longitudinal stripes, fused distally and not reaching posterior margin, and 2 lateral spots, uncovered; macrosetae reduced (holotype with only 1 brown spal s); sctl yellow pruinose, ds sctl s and sctl s long, yellow; Legs--yellow to brown; pro and meso fern light brown, brown on distal tip, meta fern pale yellow proximally, light brown distally, slightly clubbed; pro and meso tib brown, pale yellow stripe not reaching tip anteriorly, meta tib yellow in proximal half, distal half brown; first tar yellow, brown distad, remaining tar brown, setae black; emp very short, distinct; Wings--few microtrichia in proximal half, cell d with only few microtrichia along margins, densely covered with microtrichia in distal half, brown stained along veins; ptero distinct, brown; cell d terminating in 2 veins; hlt light brown. Abdomen: Black; T2-3 grey pruinose, T4-8 brown pruinose medially, anterior, lateral, and posterior margins grey pruinose, T2 anteriorly with apruinose spot, T1 with long white setae laterally, T2 in proximal third with white setae laterally, remaining T with short white and yellow setae, S grey pruinose; [male] terminalia Fig. 38--sur pointed distally, lobe dorsally and ventrally; hypd bent upwards; d aed shea tubular, medium length, v aed shea forming a plate; lat apod bifurcated. Type material--The [male] holotype is labelled 'W-Africa: Gambia: ca. 29-33 km W' Soma: Tendaba-Batelling BARKEMEYER 30.9.1987 ('30.9.' handwritten) Wegesrand (Felder, Gebusch u.a.) (handwritten on backside of label) / HOLOTYPUS Euscelidia hesperia sp. nov. det. T. Dikow 2001 (red label)'. The specimen is double mounted (minuten in block of foam), is in relatively good condition (right meta leg broken and specimen greasy; [male] terminalia attached to specimens pin in micro vial), and is deposited in the NMSA. A (2 paratype is labelled 'W-Africa: Gambia: Banjuh Bund Road (SE-Bereich) BARKEMEYER 25.9.1987 ('25.9.' handwritten) / PARATYPE Euscelidia hesperia sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is double mounted (minuten in block of foam), is in very good condition, and is deposited in the NMSA. A [female] paratype is labelled 'W-Africa: Gambia: ca. 30 km W' Soma: Duntu Malang Bolon (N' Gambia River) BARKEMEYER 28.9.1987 ('28.9.' handwritten) aufgegebene Felder bei verlassenem Dorf, z.T. feucht (handwritten on backside of label) / PARATYPE Euscelidia hesperia sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is double mounted (minuten in block of foam), is in good condition (left pped broken), and is deposited in the NMSA. [FIGURE 38 OMITTED] Type locality and distribution (Fig. 29): The Gambia, Tendaba, 13[degrees]26'N 15[degrees]49'W. The Gambia, Senegal. Material examined: SENEGAL: 4[female] [female] Tambacounda, 13[degrees]45'N 13[degrees]40'W, 6.ix. 1979 [all specimens greasy]. Depository: MRAC. Euscelidia hyalina sp. n. Figs 39, 50 Etymology: Latin adjective hyalinus = transparent; refers to the hyaline wings. Diagnosis: The species is distinguished from congeners by the predominantly apruinose sct, the apruinose T8 in the [female] [female], the brown coloration of the whole insect, and features of the [male] terminalia (sur with dorsal and ventral lobe, hypd bent upwards and divided into two lobes, d aed shea short, blunt). Description: Head: Black; fc silver pruinose, fc gib indistinct, mystax white, 6-7 macrosetae; prob and pip brownish-black, white setae; oc tr silver pruinose dorsally; occ silver pruinose, white and yellowish setae; Antennae--scp brown, brown setae ventrally; ped brown, brown setae ventrally and dorsally, white pruinose; pped brown, white pruinose; apsel brown. Thorax: Brown; ppro peg large, silver pruinose; sct brown, 3 dark brown longitudinal stripes, median stripe divided at anterior margin, lateral stripes originating medially (stripes fused distad in [male] holotype), apruinose, margins silver pruinose, white setae on pruinose area, some white setae scattered on surface; macrosetae: 1 white or black npl s, 1 black spal s; sctl silver pruinose, ds sctl s and sctl s long, brown; Legs--yellow to brown; pro and meso fem light brown, meta fern pale yellow proximally, clubbed, club brown, setae white; tib brown, pale yellow stripe not reaching tip on meta tib anteriorly; first tar proximally pale yellow, brown distad, remaining tar light brown with brown tip, black setae; emp minute; Wings--hyaline, very few microtrichia on remigium; ptero distinct, brown; cell d terminating in 3 veins; hlt light brown. Abdomen: Brown; T1 grey pruinose, T2-5 in proximal half apruinose, distal half brown pruinose, margins grey pruinose, remaining T brown pruinose, anterior and posterior margins grey pruinose, T8 in [female] apruinose, T1 with long brown setae laterally, T2 in proximal half with white setae laterally, remaining T with white setae, S2 apruinose, remaining S grey pruinose; [male] terminalia Fig. 39--sur pointed distally, lobe dorsally and ventrally; hypd bent upwards, divided into 2 lobes; d aed shea short, blunt; lat apod simple. Type material--The [male] holotype and [female] paratype are labelled '7 II 1966 [7.ii.1966] OULI BANGALA Logone, Tchad 195 Coll. J.C. Hitchcock, Jr. ('7 II' and '195' handwritten) / IN COPULA (handwritten) / HOLOTYPE [male]; PARATYPE [female] Euscelidia hyalina sp. nov. det. T. Dikow 2001 (red label)'. The specimens are double mounted (minuten in block of cork, [male] specimen above [female]), are in good condition ([male] left meta leg and left wing broken, abdomen broken (attached to block of cork), [male] terminalia attached to specimens pin in micro vial; [female] right pped broken), and are deposited in the USNM. Type locality and distribution (Fig. 50): Chad, Ouli Bangala, 7[degrees]49'N 15[degrees]49'E. Chad. [FIGURE 39 OMITTED] Euscelidia insolita sp. n. Figs 40, 41 Etymology: Latin adjective insolitus = unusual, uncommon, odd; refers to the unusual shape of the aedeagus. Diagnosis: The species is distinguished from congeners by the large size, predominantly apruinose sct, and features of the [male] terminalia (d aed shea Fig. 40). Description: Head: Black; fc yellow pruinose, fc gib indistinct, mystax white, many macrosetae; prob and plp brown, setae white; oc tr pruinose dorsally; occ silver pruinose, setae white; Antennae--scp brown, white setae ventrally, white pruinose; ped brown, brown setae ventrally and dorsally, white pruinose; pped brown, white pruinose; apsel brown. Thorax: Black; ppro peg large, silver pruinose; sct predominantly apruinose, yellow pruinosity restricted to anterior, lateral, and posterior margins, yellow setae on pruinose area, few scattered on apruinose part; macrosetae: 1 white npl s, l white or black spal s; sctl silver pruinose, ds sctl s and sctl s long, yellow; Legs--yellow to brown; fern light brown, meta fem pale yellow proximally, clubbed, club with brown lateral stripe; tib light brown, brown stripe dorsally, pale yellow stripe not reaching tip anteriorly; first tar yellow, brown distad, remaining tar brown, setae black; emp about a quarter of length of clw; Wings--densely covered with microtrichia, brown stained, veins brown; ptero distinct, brown; cell d terminating in 2 veins; hlt light brown. Abdomen: Black; grey pruinose, distal T more brown pruinose dorsally, T1 with long yellow setae laterally, T2 in proximal half with white setae laterally, remaining T with short white and yellow setae, S grey pruinose; [male] terminalia Fig. 40--sur pointed distally, lobe ventrally; hypd ventral margin straight; d aed shea proximally plate-like, distally with 4 lateral narrow lobes, dorsally with separate structure coveting ductus ejaculatotius, medium length; lat apod bifurcated. [FIGURE 40 OMITTED] Type material--The [male] holotype is labelled 'Elisabethville [Lubumbashi] Congo Beige Dr, M. Bequaert 20-I-35 [20.i. 1935] (date handwritten on backside of label) / R.I.Sc.N.B. 24.236 Coll. M. Bequaert / E. Janssens det., 1955 Euscelidia Datis Walker (handwritten except for 'E. Janssens det., 195') / HOLOTYPE Euscelidia insolita sp. nov. det. T. Dikow 2001 (red label)'. The specimen is directly mounted, is in very good condition (right meta leg broken; [male] terminalia attached to specimens pin in micro vial), and is deposited in the ISNB. Two [female] paratypes are labelled 'Elisabethville [Lubumbashi] Congo Belge Dr, M. Bequaert 20-1-35 [20.i. 1935] (respectively mars 1934 [Mar 1934]) (date handwritten on backside of label) / R.I.Sc.N.B. 24.236 Coll. M. Bequaert / PARATYPE Euscelidia insolita sp. nov. det. T. Dikow 2001 (yellow label)'. The specimens are directly mounted, are in good condition (1 specimen with left meta leg broken; 1 specimen with right meta leg and abdomen broken that is attached to specimens pin on piece of cardboard), and are deposited in the ISNB. Type locality and distribution (Fig. 41): DR Congo, Lubumbashi (former Elisabethville), 11[degrees]39'S 27[degrees]28E. DR Congo. Euscelidia Kasungu sp. n. Figs 41, 42 Etymology: None in apposition that refers to the type locality Kasungu, Malawi. [FIGURE 41 OMITTED] Diagnosis: The species is distinguished from congeners by the grey and brown pruinosity on the sct, the densely arranged microtrichia on the wings, and the pale yellow (proximal half) and brown (distal half) meta fem. Description: Head: Black; fc silver pruinose, fc gib indistinct, mystax white, many macrosetae; prob and plp brownish-black, setae white; oc tr apruinose; occ silver pruinose, setae white; Antennae--scp brown, brown setae ventrally; ped brown, brown setae ventrally and dorsally, white pruinose; pped brown, white pruinose; apsel brown. Thorax: Black; ppro peg large, silver pruinose; sct predominantly yellow pruinose, margins silver pruinose, 2 narrow median stripes, originating at anterior margin, and 2 lateral spots, not reaching anterior and posterior margins, apruinose, short yellow setae medially, white setae on margins; macrosetae: 1 white npl s, 1 black spal s; sctl silver pruinose, ds sctl s and sctl s short, white; Legs--yellow and dark brown; pro fem in proximal 2/3 yellow, distal third brown, meso fern in proximal third yellow, distal 2/3 brown, meta fern in proximal half pale yellow, clubbed, club brown, setae white; tib brown and dark brown, pale yellow stripe not reaching tip on meta tib anteriorly; first tar in proximal 2/3 pale yellow, brown distad, remaining tar brown, black setae; emp distinct, very short; Wings--cells c, sc, br, and bm with few microtrichia, remaining cells densely covered with microtrichia, less microtrichia along veins, brown stained; ptero distinct, brown; cell d terminating in 2 veins; hlt light brown. Abdomen: Black; T brown pruinose, grey pruinose laterally, T1 with long white setae laterally, T2 medially with white setae, remaining T with white and brown setae, S grey pruinose; [male] terminalia Fig. 42--sur pointed distally, lobe ventrally and slightly expanded dorsally; hypd bent upwards; d aed shea short, blunt; lat apod simple, slightly expanded proximally. Type material--The [male] holotype is labelled 'MALAWI SE1333BC 40 km S. of Kasungu 26.ii. 1987 J & A Londt Grassy hillside with rocks and bushes / HOLOTYPE Euscelidia kasungu sp. nov. det. T. Dikow 2001 (red label)'. The specimen is double mounted (minuten in block of foam), is in excellent condition ([male]; terminalia attached to specimens pin in micro vial), and is deposited in the NMSA. [FIGURE 42 OMITTED] A paratype of undeterminable gender has labels as holotype except 'PARATYPE Euscelidia kasungu sp. nov. det. T. Dikow 2001 (yellow label)'. The specimen is double mounted (minuten in block of foam), is in good condition (abdomen broken from fifth segment), and is deposited in the NMSA. Type locality and distribution (Fig. 41): Malawi, Kasungu (40 km S), 13[degrees]01'S 33[degrees]28'E (co-ordinates for Kasungu). Malawi. Euscelidia lepida sp. n. Figs 44A-D, 60 Etymology: Latin adjective lepidus = elegant, fine; refers to the beautiful, elegant appearance. Diagnosis: The species is distinguished from congeners by the broad median apruinose stripe and lateral apruinose spots on the sct, the few microtrichia on the wings, and features of the [male] terminalia (sur rounded distally, narrow ventral lobe situated proximally). Description: Head: Black; fc silver pruinose, fc gib indistinct, mystax white, 6 macrosetae; prob and pip brown-black, setae white; oc tr white pruinose dorsally; occ silver pruinose, setae white; Antennae--scp dark brown, white setae ventrally, white pruinose; ped dark brown, brown distally, brown setae ventrally and dorsally, white pruinose; pped dark brown, brown proximally, white pruinose proximally and ventrally; apsel brown. Thorax: Black; ppro peg small, distinct, silver pruinose; sct bluish-black, predominantly silver pruinose, median stripe, not reaching posterior margin, and 2 lateral spots, not reaching anterior and posterior margins, apruinose, white setae on margins; macrosetae: black, 1 npl s, 1 spal s; sctl silver pruinose, ds sctl s and sctl s short, brown: Legs--yellow to brown; fern brown, setae white, meta fem clubbed; tib brown, yellow stripe not reaching tip on meta tib anteriorly; first tar yellow, brown distad, remaining tar brown, setae black; emp little longer than half the length of clw; Wings--hyaline, microtrichia on distal margin of cell [r.sub.1] to [r.sub.5]; ptero distinct, brown; cell d terminating in 2 veins; hlt yellow. Abdomen: Black; T grey pruinose, T1 with long white and brown setae laterally, T2 in proximal half with white setae laterally, remaining T with short white and yellow setae, S grey pruinose; [male] terminalia Figs 44A-D--sur blunt, rounded distally, narrow ventral lobe proximally; hypd ventral margin straight; lat pr gonst triangular in lateral view; d aed shea short, pointed; lat apod simple. Type material--The [male] holotype is labelled 'THAILAND: Chieng Mai Province, Doi Suthep, summit 1600 m 27.ix.1981 Zool. Museum Copenhagen leg./HOLOTYPE Euscelidia lepida sp. nov. det, T. Dikow 2001 (red label).' The specimen is deposited in the ZMUC. Type locality and distribution (Fig. 60): Thailand, Chieng Mai Province, Doi Suthep, 18[degrees]48'N 98[degrees]55'E. Thailand. Euscelidia livida sp. n. Figs 43, 60 Etymology: Latin adjective lividus = black and blue; refers to the bluish-black colour of the scutum. Diagnosis: The species is distinguished from congeners by the two large lateral apruinose bluish-black spots on the sct and features of the [male] terminalia (sur with large lobe ventrally, lat apod bifurcated). Description: Head: Black; fc silver pruinose, fc gib indistinct, mystax white, many macrosetae; prob and plp brownish-black, setae white; oc tr apruinose; occ silver pruinose, setae white; Antennae (pped broken)--scp brown, brown setae ventrally; ped brown, brown setae ventrally and dorsally, white pruinose. Thorax: Orange to black; ppro peg large, silver pruinose; sct bluish-black, anterolateral and lateral margins orange, yellow pruinose medially, margins silver pruinose, 2 median narrow longitudinal stripes, not reaching posterior margin, and 2 lateral spots, not reaching anterior and posterior margins, apruinose, few white and yellow setae on margins; macrosetae: black, 1 npl s, 1 spal s; sctl silver pruinose, ds sctl s and sctl s short, white; Legs--yellow to brown; fern pale yellow proximally, light brown on remaining part, meta fern slightly clubbed, club brown; pro and meso tib light brown, pale yellow stripe anteriorly, meta rib yellow proximally, remaining part brown, pale yellow stripe not reaching tip anteriorly; first tar yellow proximally, brown distad, remaining tar brown, black setae; emp distinct, very short; Wings--hyaline, but microtrichia on distal margin, slightly browned stained; ptero distinct, brown; cell d terminating in 2 veins; hlt yellow. Abdomen: Brown; T predominantly brown pruinose, grey pruinose laterally, T1 with long brown setae laterally, T2 in proximal half with few white setae laterally, remaining T with yellow setae, S grey pruinose; [male] terminalia Fig. 43--sur pointed distally, large lobe ventrally; hypd ventral margin straight; d aed shea tubular, medium length; lat apod bifurcated. Type material--The [male] holotype is labelled 'Toungoo, Karenni 3000' [3000 ft = 900 m]. Burma IV. 14. [iv. 1914] Michelitz / HOLOTYPE Euscelidia livida sp. nov. det. T. Dikow 2001 (red label)'. The specimen is double mounted (minuten in rectangular piece of cardboard), is in very good condition (pped broken; [male] terminalia attached to specimens pin in micro vial), and is deposited in the ZMHB. Type locality and distribution (Fig. 60): Myanmar, Toungoo, 18[degrees]56'N 96[degrees]25'E. Myanmar. Euscelidia longibifida sp. n. Figs 6C, D, 7F, 8A, 14, 44E-I Etymology: Latin adjectives longus = long and bifidus = bifurcated; refers to the long lateral processes on the dorsal aedeagal sheath. Diagnosis: The species is distinguished from congeners by the yellow to brown pruinosity on the sct, the densely arranged microtrichia on the wings, and features of the [male] terminalia (d aed shea, long lat pr aed). [FIGURE 43 OMITTED] [FIGURE 44 OMITTED] Description: Head: Black; fc yellow pruinose, fc gib indistinct, mystax white, many macrosetae; prob and pip dark brown, setae white; oc tr brown pruinose dorsally; occ silver pruinose, setae white; Antennae--scp brown, black setae ventrally, white pruinose; ped light brown, brown setae ventrally and white setae dorsally, white pruinose; pped light brown, white pruinose proximally and ventrally; apsel brown. Thorax: Black; ppro peg large, silver pruinose; sct predominantly brown pruinose, margins silver pruinose, a lateral presut and a lateral psut spot may be apruinose, presut and psut dc s short, white setae on margins; macrosetae: white, 1 npl s, 1 spal s; sctl white pruinose, sctl ds and sctl s long, white; Legs--yellow to brown; fem yellow, white setae, meta fern pale yellow proximally, clubbed, club anteriorly light brown, setae short, white; pro and meso tib yellow, light brown dorsally, meta tib yellow proximally, brown distally, pale yellow stripe not reaching tip anteriorly; first tar yellow, brown distad, remaining tar brown, setae black; emp minute; Wings--densely covered with microtrichia; ptero distinct, brown; cell d terminating in 2 veins; hlt light brown. Abdomen: Black; T predominantly brown pruinose, anterior and posterior margins narrowly grey pruinose, T2 with grey pruinose ring medially, T1 with long white setae laterally, T2 in proximal third with white setae laterally, remaining T with short white and brown setae, S grey pruinose; [male] terminalia Figs 44E-I--sur pointed distally, lobe ventrally; hypd bent upwards, divided into 2 separated lobes; d aed shea short, lat pr aed long, extending gonp; lat apod simple. Type material--The [male] holotype is labelled 'SOUTH AFRICA 2427BA Transv. [Transvaal] Bulge River 30.I.1978 JGH. Londt Open grassveld area / HOLOTYPE Euscelidia longibifida sp. nov. det. T. Dikow 2001 (red label)'. The specimen is double mounted (minuten in rectangular piece of cardboard), is in excellent condition, and is deposited in the NMSA. The 6 [female] [female] 5[male][male] paratypes have same labels as holotype except 'PARATYPE Euscelidia longibifida sp. nov. det. T. Dikow 2001 (yellow label)'. The specimens are double mounted (minuten in rectangular piece of cardboard), are in good to very good condition, and are deposited in the NMSA. Type locality and distribution (Fig. 14): South Africa, Northern Province, Bulge River, 24[degrees]06'S 27[degrees]41'E. Mozambique, South Africa. Material examined: MOZAMBIQUE: 1[male] Henrique, 15[degrees]18'S 39[degrees]58'E, 15.xii.; SOUTH AFRICA: KwaZulu-Natal: 1? Kube Yini Game Reserve, 27[degrees]28'S 32[degrees]14'E, 10-14.i.1994; 2 [female] [female] 1[male] 3?? Mfongosi, 27[degrees]17'S 32[degrees]09'E, iii.1914, ii., iii.1917; 1[female] Weenen, 28[degrees]51'S 30[degrees]05'E, i.1924; 1[male] 1? Ashburton, 29[degrees]38'S 30[degrees]24'E, 2.i.1989; 1[male] 1? Mackston, 30[degrees]01'S 30[degrees]18'E, iv.1977; Northern Prov: 5 [female] [female] 2[male][male] Potgietersrus, 24[degrees]15'S 28[degrees]58'E, 28.i.1978; 5 [female] [female] 2[male][male] Northam, 24[degrees]56'S 27[degrees]15'E, 1.xi.1978; 1[male] Warmbaths, 24[degrees]52'S 28[degrees]17'E, 1.xi.1978; 1 [female] Thabazimbi, 24[degrees]35'S 27[degrees]24'E, 1.xi.1978; North-West Prov: 3 [female] [female] 4[male][male] Brits, 2527DA, 2.iii.1978. Depository: BMNH, NMSA. Euscelidia lucida Oldroyd, 1939 Figs 45A-D, 46 Euscelidia lucida Oldroyd, 1939: 28; Hull 1962: 305; Oldroyd 1980: 356. Diagnosis: The species is distinguished from congeners by the predominantly apruinose sct and features of the [male] terminalia (Figs 45A-D). Redescription: Head: Black; fc silver pruinose, fc gib indistinct, mystax white, 5-8 macrosetae; oc tr white pruinose dorsally; occ yellow pruinose, few white setae; Antennae--scp and ped light brown, setae brown; pped yellow, white pruinose; apsel hyaline; Thorax: Orange to brown; sct brown, predominantly yellow pruinose, a median longitudinal stripe, not reaching posterior margin, and 2 lateral longitudinal stripes, not reaching anterior margin, apruinose, surface of stripes black darkened, lateral and posterior margins with few yellow setae, macrosetae: black, 1 npl s, 1 spal s; sctl silver pruinose, ds sctl s and sctl long, yellow; Legs--yellow and light brown; pro and meso fern light brown, proximal half yellow posteriorly, meta fern pale yellow proximally, light brown, slightly clubbed, club with brown stripe laterally, setae white; tib light brown, yellow stripe not reaching tip on meta tib anteriorly; emp distinct, very short (emp longer in holotype); Wings--hyaline, some microtrichia distad; ptero distinct, brown; cell d terminating in 2 veins; hlt yellow with brown knob; Abdomen: Brown; T predominantly brown pruinose, apruinose spot on T2-5 anteriorly, posterior margin grey pruinose, T1 with brown setae laterally, T2 with few white setae laterally, remaining T with short yellow setae, S grey pruinose; [male] terminalia Figs 45A-D--sur pointed distally, lobe dorsally and ventrally, dorsal lobe larger and pointed; hypd bent upwards; d aed shea short, blunt; lat apod simple. [FIGURE 45 OMITTED] Type material--The [male] holotype is labelled 'Type [male] (circular label with red submarginal border) Euscelidia lucida Oldr. (on backside) / Namwamba Valley. 6,500 ft. [1950 m] F. Edwards. / UGANDA: Ruwenzori Range. xii.1934-i.1935. B.M.E. Afr. Exp. [British Museum East Africa Expedition] B.M. 1935-203 / Euscelidia lucida sp. nov. H. Oldroyd, det. 1938. (species name handwritten) / HOLOTYPE Euscelidia lucida Oldroyd det. J.E. Chainey 1982 (handwritten) Holotype (circular label with red border, attached to the other label)'. The specimen is double mounted (minuten in rectangular piece of plastic), is in good condition (abdomen broken ([male] terminalia were attached to piece of plastic); [male] terminalia attached to specimens pin in micro vial), and is deposited in the BMNH. Type locality and distribution (Fig. 46): Uganda, Ruwenzori Range, Namwamba Valley, 0[degrees]05'N 30[degrees]10'E. DR Congo, Nigeria, Uganda. Material examined: DR CONGO: 1 [female] Virunga National Park, Tshiaberimu, 0[degrees]08'S 29[degrees]25'E, 29.iii.1954; NIGERIA: 1 [female] Ngel Nyaki, 7[degrees]04'N l l[degrees]03'E, 28.xi.-3.xii.1968. Depository: BMNH, MRAC. Euscelidia lucioides sp. n. Figs 45E-I, 46 Etymology: Refers to the similarity of this species to lucida. Diagnosis: The species is distinguished from congeners by the predominantly grey pruinose sct, a brown apruinose median stripe and two apruinose lateral stripes on the sct, and features of the [male] terminalia. [FIGURE 46 OMITTED] Description: Head: Black; fc silver pruinose (in upper half more yellow pruinose), fc gib indistinct, mystax white, 6-8 macrosetae; prob and plp brown, setae white; oc tr white pruinose dorsally; occ silver pruinose, few white and yellowish setae; Antennae--scp yellow, white setae ventrally, white pruinose; ped yellow, brown setae ventrally and dorsally, white pruinose; pped yellow, brown distad, white pruinose; apsel brown. [FIGURE 47 OMITTED] Thorax: Brown; ppro peg small, distinct, silver pruinose; sct predominantly yellow pruinose, 3 apruinose longitudinal stripes, median stripe not reaching posterior margin, lateral ones not reaching anterior and posterior margins, yellow setae on margins; macrosetae: brown, 1 npl s, 1 spal s; sctl white pruinose, ds sctl s and sctl s long, light brown; Legs--yellow to brown; fem light brown, yellow proximally, meta fern slightly clubbed; pro and meso tib light brown, pale yellow stripe anteriorly, meta tib brown, pale yellow stripe not reaching tip anteriorly; tar light brown, brown distad, black setae; emp minute; Wings--hyaline, few microtrichia scattered on remigium and on distal margin, slightly brown stained; ptero distinct, brown; cell d terminating in 2 veins; hlt light brown. Abdomen: Brown; T predominantly brown pruinose, anterior and posterior margins narrowly grey pruinose, T2 with apruinose spot medially, T3 with apruinose spot proximally, T1 with long brown setae laterally, T2 in proximal half with white setae laterally, remaining T with white to brown setae, S grey pruinose, S2 in proximal half apruinose; [male] terminalia Figs 45E-I--sur pointed distally, lobe dorsally and ventrally; hypd bent upwards; d aed shea short, blunt; lat apod simple. Type material--The [male] holotype is labelled 'KENYA, Ngong Forestry Station 1-7 Feb. 1968 [1-7.ii.1962] Malaise trap Krombein & Spangler / HOLOTYPE Euscelidia lucioides sp. nov. det. T. Dikow 2001 (red label)'. The specimen is directly mounted, is in excellent condition (many Lepidoptera scales on specimen), and is deposited in the USNM. The 2[female] [female] 2[male][male] paratypes are labelled 'KENYA, Ngong Forestry Station 13-18 Jan. [1[male]] 27-31 Jan. [2[female] [female]] 1-7 Feb. 1968 [1[male]] Malaise trap Krombein & Spangler / PARATYPE Euscelidia lucioides sp. nov. det. T. Dikow 2001 (yellow label)'. The specimens are directly mounted, are in very good to excellent condition (many Lepidoptera scales on specimens), and are deposited in the USNM. Type locality and distribution (Fig. 46): Kenya, Ngong, 1[degrees]21'S 36[degrees]39'E. Kenya. Remarks: The species is very similar to lucida. Some features of the [male] terminalia (shape of sur and aed) as well as the shorter empodium, however, support my decision in describing it as a different species.
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